A new species of highly troglobitic cockroach, Nocticola baumi n. sp., is described from limestone caves of Papua Province, Indonesia. The new species is the first Nocticola reported in New Guinea island and differs from other known Nocticola spp. by the combination of the following characters: compound eyes and ocelli absent, tegmina fully developed extending beyond the abdomen, hind wings absent, legs extremely long. The discovery of the genus on New Guinea bridges the gap between Oriental and Australian distribution of the genus and stresses the unknown diversity of the genus.

Department of Zoology and Fisheries Faculty of Agrobiology Food and Natural Resources Czech University of Life Sciences Prague Prague Czech Republic

Entomology Section Museum of Natural History University of the Philippines Los Baos Philippines

Museum Zoologicum Bogoriense Research Center for Biology Indonesian Institute of Sciences Jakarta Indonesia

South Bohemian Research Center of Aquaculture and Biodiversity of Hydrocenoses Faculty of Fisheries and Protection of Waters University of South Bohemia in esk Budjovice Vodany Czech Republic

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Dating in the Dark: Elevated Substitution Rates in Cave Cockroaches (Blattodea: Nocticolidae) Have Negative Impacts on Molecular Date Estimates