The Mediterranean is characterised by high biodiversity and numerous endemic species. These species are not only present in natural habitats, but also inhabit areas under human influence, such as agricultural lands. In the biodiversity assessment of Mediterranean vineyards and olive orchards within Zadar County, in Croatia, we identified eight endemic species with Mediterranean distribution, six with a Balkan Peninsula distribution, four with Dinaric Alps distribution and three species rare and endangered in Europe. Alongside these species, we have recorded five new species for Croatian fauna, many of those identified by combining morphological characteristics and the DNA barcoding tool. Araneae and Coleoptera contributed the highest number of endemic species and groups with new record were the following: Coleoptera, Diptera and Araneae. Compared to other sites, an olive orchard with ecological pest management (EPM), surrounded by natural ecosystems, had the highest ratio of endemic and rare species. Our findings emphasise that agricultural lands in the Mediterranean can be habitats for endemic and rare species and that future biodiversity research of these habitats is highly important, to monitor potential biodiversity changes and motivate future species and ecosystem conservation.

Croatian Natural History Museum Demetrova 1 Zagreb Croatia Croatian Natural History Museum Demetrova 1 Zagreb Croatia

Department for Ecology Agronomy and Aquaculture University of Zadar Trg kneza Višeslava Zadar Croatia Department for Ecology Agronomy and Aquaculture University of Zadar Trg kneza Višeslava Zadar Croatia

Department of Biology Faculty of Science University of Zagreb Rooseveltov trg 6 Zagreb Croatia Department of Biology Faculty of Science University of Zagreb Rooseveltov trg 6 Zagreb Croatia

Department of Botany and Zoology Faculty of Science Masaryk University Kotlarska 2 Brno Czech Republic Department of Botany and Zoology Faculty of Science Masaryk University Kotlarska 2 Brno Czech Republic

Varaždin City Museum Šetalište Josipa Jurja Strossmayera 3 Varaždin Croatia Varaždin City Museum Šetalište Josipa Jurja Strossmayera 3 Varaždin Croatia

Zobrazit více v PubMed

Allegrucci Giuliana, Rampini Mauro, Di Russo Claudio, Lana Enrico, Cocchi Sara, Sbordoni Valerio. Phylogeography and systematics of the westernmost Italian Dolichopoda species (Orthoptera, Rhaphidophoridae) ZooKeys. 2014;437:1–23. doi: 10.3897/zookeys.437.7917. PubMed DOI PMC

Anđelić Dmitrović Barbara, Jelić Mišel, Rota Emilia, Šerić Jelaska Lucija. DNA barcoding of invertebrates inhabiting olive orchards and vineyards accelerates understudied Mediterranean biodiversity assessment. Diversity. 2022;14(3) doi: 10.3390/d14030182. DOI

Anichtchenko Alexandre, Guéorguiev Borislav. Taxonomic notes on Pelor Bonelli, with description of a new species from Greece (Coleoptera: Carabidae: Zabrini) Biologia. 2009;64(5):937–941. doi: 10.2478/s11756-009-0144-1. DOI

Bahlai Christine A., Xue Yingen, McCreary Cara M., Schaafsma Arthur W., Hallett Rebecca H. Choosing organic pesticides over synthetic pesticides may not effectively mitigate environmental risk in soybeans. PLOS One. 2010;5(6) doi: 10.1371/journal.pone.0011250. PubMed DOI PMC

Baker Brian P., Green Thomas A., Loker Ali J. Biological control and integrated pest management in organic and conventional systems. Biological Control. 2020;140:e11250. doi: 10.1016/j.biocontrol.2019.104095. DOI

Bank R. A., Neubert E. Checklist of the land and freshwater Gastropoda of Europe. Last update. 2020;July 16th, 2017

Barić B., Pajač Živković I. Suzbijanje prezimljujuće populacije jabukova savijača primjenom entomopatogenih nematoda. Fragmenta Phytomedica. 2020;34(5):23–31.

Boer Peter. Revision of the European ants of the Aphaenogastertestaceopilosa-group (Hymenoptera: Formicidae) Tijdschrift voor Entomologie. 2013;156(1):57–93. doi: 10.1163/22119434-00002022. DOI

Borowiec L., Salata S. Ants of Greece-checklist, comments and new faunistic data (Hymenoptera: Formicidae. Genus. 2012;23(4):461–563.

Borowiec L. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae) Genus. 2014;25(1/2):1–340.

Bosmans R. Revision of the genus Zodarion Walckenaer, 1833, part II Western and Central Europe, including Italy (Araneae: Zodariidae) Bulletin-British Arachnological Society. 1997;10:265–294.

Bourgoin, T FLOW (Fulgoromorpha Lists on The Web): a world knowledge base dedicated to Fulgoromorpha. http://hemiptera-databases.org/flow/ [2022-05-12T00:00:00+03:00]. http://hemiptera-databases.org/flow/

Bračko, G Review of the ant fauna (Hymenoptera: Formicidae) of Croatia. Acta Entomologica Slovenica. 2006;14:2.

Brandmayr P., Brandmayr Zetto T. Parental behaviour and aspects of the biology of Carterus (Sabienus) calydonius Rossi and considerations on parental behaviour already reported on Carabids (Coleoptera, Carabidae) Redia. 1974;55:143–175.

Brock P. D. A photographic guide to Insects of Southern Europe & the Mediterranean. Pisces Publications; 2017.

Science California Academy of. AntWeb. Version 8.42. https://www.antweb.org. [2022-03-23T00:00:00+02:00]. https://www.antweb.org.

Canestrini G., Pavesi P. Araneidi italiani. Atti della Società Italiana di Scienze Naturali. 1868;11(3):738–872.

Cardenas M., Pascual F., Campos M., Pekar S. The spider assemblage of olive groves under three management systems. Environmental Entomology. 2015;44(3):509–518. doi: 10.1093/ee/nvv030. PubMed DOI

Cárdenas Manuel, Ruano Francisca, García Pedro, Pascual Felipe, Campos Mercedes. Impact of agricultural management on spider populations in the canopy of olive trees. Biological Control. 2006;38(2):188–195. doi: 10.1016/j.biocontrol.2006.02.004. DOI

Cardoso Pedro, Henriques Sérgio S., Gaspar Clara, Crespo Luis C., Carvalho Rui, Schmidt Jesper B., Sousa Pedro, Szűts Tamás. Species richness and composition assessment of spiders in a Mediterranean scrubland. Journal of Insect Conservation. 2007;13(1):45–55. doi: 10.1007/s10841-007-9116-3. DOI

Chimeno Caroline, Hausmann Axel, Schmidt Stefan, Raupach Michael J., Doczkal Dieter, Baranov Viktor, Hübner Jeremy, Höcherl Amelie, Albrecht Rosa, Jaschhof Mathias, Haszprunar Gerhard, Hebert Paul D. N. Peering into the darkness: DNA barcoding reveals surprisingly high diversity of unknown species of Diptera (Insecta) in Germany. Insects. 2022;13(1) doi: 10.3390/insects13010082. PubMed DOI PMC

Chukwuka Christian O., Ejere Vincent C., Asogwa Chinweike N., Nnamonu Emmanuel I., Okeke Ogochukwu C., Odii Elijah I., Ugwu Godwin C., Okanya Loretta C., Levi Chidinma A. Eco-physiological adaptation of the land snail Achatinaachatina (Gastropoda: Pulmonata) in tropical agro-ecosystem. The Journal of Basic & Applied Zoology. 2014;67(2):48–57. doi: 10.1016/j.jobaz.2014.06.001. DOI

Cigliano M. M., Braun H, Eades D. C, Otte D. Orthoptera Species File. Version 5.0/5.0. http://Orthoptera.SpeciesFile.org http://Orthoptera.SpeciesFile.org

Ciplak B., Willemse F., Chobanov D., Heller K. G. Systematic status and distributi-on of Eupholidoptera (Orthoptera: Tettigoniidae) in the Balkans (north of Central Greece) Articulata . 2007;22(1):33–46.

Cuff Jordan P., Drake Lorna E., Tercel Maximillian P. T.G., Stockdale Jennifer E., Orozco‐terWengel Pablo, Bell James R., Vaughan Ian P., Müller Carsten T., Symondson William O. C. Money spider dietary choice in pre‐ and post‐harvest cereal crops using metabarcoding. Ecological Entomology. 2021;46(2):249–261. doi: 10.1111/een.12957. DOI

Cuttelod A., Seddon M., Neubert E. European red list of non-marine molluscs. Publications Office of the European Union; 2011.

de Jong Yde, Kouwenberg Juliana, Boumans Louis, Hussey Charles, Hyam Roger, Nicolson Nicola, Kirk Paul, Paton Alan, Michel Ellinor, Guiry Michael, Boegh Phillip, Pedersen Henrik, Enghoff Henrik, von Raab-Straube Eckhard, Güntsch Anton, Geoffroy Marc, Müller Andreas, Kohlbecker Andreas, Berendsohn Walter, Appeltans Ward, Arvanitidis Christos, Vanhoorne Bart, Declerck Joram, Vandepitte Leen, Hernandez Francisco, Nash Róisín, Costello Mark, Ouvrard David, Bezard-Falgas Pascale, Bourgoin Thierry, Wetzel Florian, Glöckler Falko, Korb Günther, Ring Caroline, Hagedorn Gregor, Häuser Christoph, Aktaç Nihat, Asan Ahmet, Ardelean Adorian, Borges Paulo, Dhora Dhimiter, Khachatryan Hasmik, Malicky Michael, Ibrahimov Shaig, Tuzikov Alexander, De Wever Aaike, Moncheva Snejana, Spassov Nikolai, Chobot Karel, Popov Alexi, Boršić Igor, Sfenthourakis Spyros, Kõljalg Urmas, Uotila Pertti, Gargominy Olivier, Dauvin Jean-Claude, Tarkhnishvili David, Chaladze Giorgi, Tuerkay Michael, Legakis Anastasios, Peregovits László, Gudmundsson Gudmundur, Ólafsson Erling, Lysaght Liam, Galil Bella, Raimondo Francesco, Domina Gianniantonio, Stoch Fabio, Minelli Alessandro, Spungis Voldermars, Budrys Eduardas, Olenin Sergej, Turpel Armand, Walisch Tania, Krpach Vladimir, Gambin Marie, Ungureanu Laurentia, Karaman Gordan, Kleukers Roy, Stur Elisabeth, Aagaard Kaare, Valland Nils, Moen Toril, Bogdanowicz Wieslaw, Tykarski Piotr, Węsławski Jan, Kędra Monika, M. de Frias Martins Antonio, Abreu António, Silva Ricardo, Medvedev Sergei, Ryss Alexander, Šimić Smiljka, Marhold Karol, Stloukal Eduard, Tome Davorin, Ramos Marian, Valdés Benito, Pina Francisco, Kullander Sven, Telenius Anders, Gonseth Yves, Tschudin Pascal, Sergeyeva Oleksandra, Vladymyrov Volodymyr, Rizun Volodymyr, Raper Chris, Lear Dan, Stoev Pavel, Penev Lyubomir, Rubio Ana, Backeljau Thierry, Saarenmaa Hannu, Ulenberg Sandrine. PESI - a taxonomic backbone for Europe. Biodiversity Data Journal. 2015;3 doi: 10.3897/bdj.3.e5848. PubMed DOI PMC

De Jong Y., Verbeek M., Michelsen V., Bjørn de Place, Los W., Steeman F., Penev L., et al. Fauna Europaea-all European animal species on the web. Biodiversity Data Journal. 2014;2 PubMed PMC

Deltshev C. A faunistic and zoogeographical review of the spiders (Araneae) of the Balkan Peninsula. Journal of Arachnology. 1999:255–261.

Deltshev C., Naumova M., Matevski D., Indzhov S. New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species. Zootaxa. 2022;5174(3):247–261. doi: 10.11646/zootaxa.5174.3.3. PubMed DOI

De Mattia Willy, Pešic Vladimir. Taxonomic and nomenclatural notes on Dalmatian and Montenegrin Tandonia: old issues solved and new problems arise (Gastropoda: Pulmonata: Milacidae) Folia Malacologica. 2015;23(3):197–210. doi: 10.12657/folmal.023.017. DOI

Di Franco F. New considerations about the gnaphosid fauna of Italy (Araneae: Gnaphosidae. Bulettin-British Arachnological Society. 1997;10:242–246.

Drakšić M., Katušić L. Preliminary results on the spider (Araneae) fauna of NP Kornati (Croatia); Entomologica Croatica; XXII Symposium Internationale Entomofaunisticum Europae Centralis; Varaždin, Croatia. 2011. 13

Duque-Trujillo D., Hincapié C. A., Osorio M., Zartha-Sossa J. W. Strategies for the attraction and conservation of natural pollinators in agroecosystems: a systematic review. International Journal of Environmental Science and Technology. 2022 doi: 10.1007/s13762-022-04634-6. DOI

Evenhuis N. L., Pape Thomas, Pont A. C. Nomenclatural studies toward a world list of diptera genus-group names. Part V: Pierre-Justin-Marie Macquart. Zootaxa. 2016;4172(1) doi: 10.11646/zootaxa.4172.1.1. PubMed DOI

Francuski Ljubinka, Ludoški Jasmina, Vujić Ante, Milankov Vesna. Phenotypic evidence for hidden biodiversity in the Merodonaureus group (Diptera, Syrphidae) on the Balkan Peninsula: conservation implication. Journal of Insect Conservation. 2011;15(3):379–388. doi: 10.1007/s10841-010-9311-5. DOI

Franin K., Franin G. K., Maricic B., Marcelic S., Pavlovic M., Kos T., Laznik Z. True bugs (Heteroptera) assemblage and diversity in the ecological infrastructures around the Mediterranean vineyards. Bulletin of Insectology. 2021;74:65–78.

Freude H., Harde K. W., Lohse G. A., Klausnitzer B. Die Käfer Mitteleuropas. Band 2. Adephaga. 1. Carabidae (Laufkäfer) Springe Spektrum; 2004.

Froidevaux Jérémy S. P., Louboutin Bastien, Jones Gareth. Does organic farming enhance biodiversity in Mediterranean vineyards? A case study with bats and arachnids. Agriculture, Ecosystems & Environment. 2017;249:112–122. doi: 10.1016/j.agee.2017.08.012. PubMed DOI PMC

Gajski Domagoj, Pekár Stano. Assessment of the biocontrol potential of natural enemies against psyllid populations in a pear tree orchard during spring. Pest Management Science. 2021;77(5):2358–2366. doi: 10.1002/ps.6262. PubMed DOI

Gasperini R. Prilog fauni dalmatinskih pauka (Araneae et Opiliones) Godisnij Izvejestaj vel. Real- ke Splitu . 1891:1–18.

org. GBIF. GBIF.org: GBIF Home Page. https://www.gbif.org https://www.gbif.org

Gnezdilov V. M., Holzinger W. E., Wilson M. R. The Western Palaearctic Issidae (Hemiptera, Fulgoroidea): an illustrated checklist and key to genera and subgenera. Trudy Zoologičeskogo Instituta RAN. 2014;318(S1):1–136.

Godan D. Pest slugs and snails. Biology and control. Springer; Berlin: 1983. DOI

Goidanich A. Studio delle forme liburniche del genere Carabus (Coleoptera Adephaga) e della loro distribuzione. Bollettino del Laboratorio di Entomologia Bologna. 1932;5:53–84.

Grbac I., Katušić L., Lukić M. Catalogue of spiders (Araneae) deposited in the Croatian Natural History Museum. Natura Croatica. 2019;28(1)

Griffiths H. I., Krystufek B., Reed J. M. Balkan biodiversity. Vol. 10. Kluwer Academic Publishers; 2004. DOI

Gunstone Tari, Cornelisse Tara, Klein Kendra, Dubey Aditi, Donley Nathan. Pesticides and soil invertebrates: A hazard assessment. Frontiers in Environmental Science. 2021;9 doi: 10.3389/fenvs.2021.643847. DOI

Hammer Ø., Harper D. A., Ryan P. D. PAST: Paleontological statistics software package for education and data analysis. Palaeontologia Electronica. 2001;4(1):9.

Happe Anne-Kathrin, Alins Georgina, Blüthgen Nico, Boreux Virginie, Bosch Jordi, García Daniel, Hambäck Peter A., Klein Alexandra-Maria, Martínez-Sastre Rodrigo, Miñarro Marcos, Müller Ann-Kathrin, Porcel Mario, Rodrigo Anselm, Roquer-Beni Laura, Samnegård Ulrika, Tasin Marco, Mody Karsten. Predatory arthropods in apple orchards across Europe: Responses to agricultural management, adjacent habitat, landscape composition and country. Agriculture, Ecosystems & Environment. 2019;273:141–150. doi: 10.1016/j.agee.2018.12.012. DOI

Harz Kurt. Die Orthopteren Europas / The Orthoptera of Europe. The Hague; 1969. DOI

Hippa Heikki, Vilkamaa Pekka, Heller Kai. Review of the Holarctic Corynoptera Winnertz, 1867, s. str. (Diptera, Sciaridae) Zootaxa. 2010;2695(1) doi: 10.11646/zootaxa.2695.1.1. PubMed DOI

Hochkirch A., et al. European red list of grasshoppers, crickets and bush-crickets. Publications Office of the EU 2016

Hurka K. Carabidae of the Czech and Slovak republics. Print-centrum; 1996.

Ichihara Minoru, Matsuno Kazuo, Inagaki Hidehiro, Saiki Chieko, Mizumoto Shunsuke, Yamaguchi Shou, Yamashita Masayuki, Sawada Hitoshi. Creation of paddy levees to enhance the ecosystem service of weed seed predation by crickets. Landscape and Ecological Engineering. 2015;11(1):227–233. doi: 10.1007/s11355-014-0254-y. DOI

Ivanković Tatalović Lara, Anđelić Barbara, Jelić Mišel, Kos Tomislav, A. Benítez Hugo, Šerić Jelaska Lucija. Fluctuating asymmetry as a method of assessing environmental stress in two predatory carabid species within Mediterranean agroecosystems. Symmetry. 2020;12(11) doi: 10.3390/sym12111890. DOI

Jambrošić Vladić Željka, Benítez Hugo A., Pirnat Alja, Hristovski Slavčo, Jelaska Lucija Šerić. Variations in body shape of mountain habitat specialist Carabuscroaticus and its sister species Carabuscaelatus (Coleoptera: Carabidae) populations across Dinaric Alps. Zoomorphology. 2019;138(1):85–96. doi: 10.1007/s00435-018-0428-5. DOI

Jambrošić Vladić Željka. Ekologija i biogeografija odabranih endemskih epigejskih vrsta trčaka (Coleoptera: Carabidae) dinarskog krša. University of Zagreb. Faculty of Science. Department of Biology; 2020.

Jambrošić Vladić Željka, Šerić Jelaska Lucija. Long term changes (1990-2016) in carabid beetle assemblages (Coleoptera: Carabidae) in protected forests on Dinaric Karst on Mountain Risnjak, Croatia. European Journal of Entomology. 2020;117:56–67. doi: 10.14411/eje.2020.006. DOI

Jantscher E. Diagnostic characters of Xysticuscristatus, X.audax and X.macedonicus (Araneae: Thomisidae) Bulletin-British Arachnological Society. 2001;12(1):17–25.

Jelaska Sven D., Nikolić Toni, Šerić Jelaska Lucija, Kušan Vladimir, Peternel Hrvoje, Gužvica Goran, Major Zoran. Terrestrial biodiversity analyses in Dalmatia (Croatia): A complementary approach using diversity and rarity. Environmental Management. 2010;45(3):616–625. doi: 10.1007/s00267-010-9437-y. PubMed DOI

Katušić Luka. Pauci Hrvatske i analiza okolišne uvjetovanosti njihove rasprostranjenosti. University of Zagreb. Faculty of Science. Department of Biology; 2017.

Kavvadias V., Koubouris G. In: Soil Fertility Management for Sustainable Development. Panpatte D., Jhala Y., editors. Springer; Singapore: 2019. Sustainable soil management practices in olive groves. DOI

Kenyeres Zoltán, Rácz István A., Varga Zoltán. Endemism hot spots, core areas and disjunctions in European Orthoptera. Acta Zoologica Cracoviensia - Series B: Invertebrata. 2009;52(1):189–211. doi: 10.3409/azc.52b_1-2.189-211. DOI

Komnenov M. Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld. 2010;5(2009):52–69.

Kromp Bernhard. Carabid beetles in sustainable agriculture: a review on pest control efficacy, cultivation impacts and enhancement. Agriculture, Ecosystems & Environment. 1999;74:187–228. doi: 10.1016/s0167-8809(99)00037-7. DOI

Kuntner M. A contribution to the knowledge of the Slovenian spider fauna: eleven species new for Slovenia and some other interesting findings (Arachnida, Araneae); Proceedings of the 16th European Colloquium of Arachnology; 16th European Colloquium of Arachnology; Siedlce, Slovenia. 1997. 8

Kurbalija Ana. Pregled entomofaune močvarnih staništa od međunarodnog značenja u Republici Hrvatskoj. Odjel za biologiju, Osijek; 2012.

Lebas C., Galkowski C., Blatrix R., Wegnez P. Ants of Britain and Europe. Bloomsbury Publishing PLC; London: 2019.

Löbl L., Löbl D. Archostemata-Myxophaga-Adephaga. Revised and Updated. Brill Academic Publishers; Leiden, Netherlands: 2017. DOI

Macfadyen A. Notes on methods for the extraction of small soil arthropods. The Journal of Animal Ecology. 1953:65–77. doi: 10.2307/1691. DOI

Mathis Wayne N., Zatwarnicki Tadeusz. New Species and Taxonomic Clarifications for Shore Flies from the Delmarva States (Diptera: Ephydridae) Proceedings of the Entomological Society of Washington. 2010;112(1):97–128. doi: 10.4289/0013-8797-112.1.97. DOI

Melichar L. Monographie der Issiden (Homoptera) National Agricultural Library; Maryland, USA: 1906.

Eds. MolluscaBase., editor. MolluscaBase. https://www.molluscabase.org https://www.molluscabase.org

Moreby S. J., Sotherton N. W., Jepson P. C. The effects of pesticides on species of non‐target Heteroptera inhabiting cereal fields in southern England. Pesticide Science. 1997;51(1):39–48. doi: 10.1002/(SICI)1096-9063(199709)51:1<39::AID-PS611>3.0.CO;2-3. DOI

Morinière Jérôme, Balke Michael, Doczkal Dieter, Geiger Matthias F., Hardulak Laura A., Haszprunar Gerhard, Hausmann Axel, Hendrich Lars, Regalado Ledis, Rulik Björn, Schmidt Stefan, Wägele Johann‐Wolfgang, Hebert Paul D. N. A DNA barcode library for 5,200 German flies and midges (Insecta: Diptera) and its implications for metabarcoding‐based biomonitoring. Molecular Ecology Resources. 2019;19(4):900–928. doi: 10.1111/1755-0998.13022. PubMed DOI PMC

Muster C., Bosmans R., Thaler K. The Philodromus pulchellus-group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae) Invertebrate Systematics. 2007;21(1):39–72. doi: 10.1071/IS06014. DOI

Nentwig W., et al. Spiders of Europe. https://www.araneae.nmbe.ch. [2022-03-23T00:00:00+02:00]. https://www.araneae.nmbe.ch

Nyffeler M., Sunderland K. D. Composition, abundance and pest control potential of spider communities in agroecosystems: a comparison of European and US studies. Agriculture, Ecosystems & Environment. 2003;95(2-3):579–612. doi: 10.1016/S0167-8809(02)00181-0. DOI

Özdikmen H., Bal N., Coral Şahin D. New flea beetles records of Aphthona Chevrolat in Turkey (Chrysomelidae: Galerucinae: Alticini. Munis Entomology & Zoology. 2018;13(2):395–400.

Pantini Paolo, Isaia Marco. Araneae.it: the online Catalog of Italian spiders, with addenda on other Arachnid Orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae) Fragmenta Entomologica. 2019;51(2):127–152. doi: 10.4081/fe.2019.374. DOI

Picchi Malayka Samantha. Spiders (Araneae) of olive groves and adjacent semi-natural habitats from central Italy. Arachnologische Mitteilungen: Arachnology Letters. 2020;60(1):1–11. doi: 10.30963/aramit6001. DOI

Pintilioaie Alexandru-Mihai, Spaseni Petronel, Jurjescu Ana, Rădac Ioan Alexandru. First record of the alien mantid Hierodulatenuidentata (Insecta: Mantodea) in Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”. 2021;64(1):37–49. doi: 10.3897/travaux.64.e65489. DOI

Pisa L. W., Amaral-Rogers V., Belzunces L. P., Bonmatin J. M., Downs C. A., Goulson D., Kreutzweiser D. P., Krupke C., Liess M., McField M., Morrissey C. A., Noome D. A., Settele J., Simon-Delso N., Stark J. D., Van der Sluijs J. P., Van Dyck H., Wiemers M. Effects of neonicotinoids and fipronil on non-target invertebrates. Environmental Science and Pollution Research. 2015;22(1):68–102. doi: 10.1007/s11356-014-3471-x. PubMed DOI PMC

Previšić Ana, Graf Wolfram, Vitecek Simon, Kučinić Mladen, Bálint Miklós, Keresztes Lujza, Pauls Steffen U, Waringer Johann. Cryptic diversity of caddisflies in the Balkans: the curious case of Ecclisopteryx species (Trichoptera: Limnephilidae) Arthropod Systematics & Phylogeny. 2014;72(3):309–329. PubMed PMC

Ratnasingham Sujeevan, Hebert Paul D. N. BARCODING: bold: The Barcode of Life Data System (http://www.barcodinglife.org) Molecular Ecology Notes. 2007;7(3):355–364. doi: 10.1111/j.1471-8286.2007.01678.x. PubMed DOI PMC

Ratnasingham Sujeevan, Hebert Paul D. N. A DNA-based registry for all animal species: The Barcode Index Number (BIN) System. PLOS One. 2013;8(7) doi: 10.1371/journal.pone.0066213. PubMed DOI PMC

Ricketts Taylor, Imhoff Marc. Biodiversity, urban areas, and agriculture: Locating priority ecoregions for conservation. Conservation Ecology. 2003;8(2) doi: 10.5751/es-00593-080201. DOI

Rukavina I., Mrazović A, Kučinić M., Šerić Jelaska L. Assemblage, zoogeography and endangered status of carabid beetles in forest habitats of the Učka Nature Park. Entomologia Croatica. 2010;14(1-2):121–134.

Samiayyan K. Spiders – The generalist super predators in agro-ecosystems. Integrated Pest Management. 2014:283–310. doi: 10.1016/b978-0-12-398529-3.00016-6. DOI

Schowalter Timothy, Chao Jung-Tai. Canopy Insect Sampling. Measuring Arthropod Biodiversity. 2021:467–493. doi: 10.1007/978-3-030-53226-0_18. DOI

Seifert B. The ants of central and north Europe. lutra Verlags-und Vertriebsgesellschaft; 2018.

Šerić Jelaska L., Vujčić-Karlo S, Durbešić P. Notes on the taxonomy of the genus Carabus L. (Coleoptera: Carabidae) in Croatia. Acta entomologica Slovenica. Ljubljana. 2004;12:129–138.

Šerić Jelaska L., Temunović M. Assessing the conservation status of lower course of Una river using records of Adephagan Coleoptera , editor. 9th European Congress of Entomology 2010

Šerić Jelaska Lucija, Franjević Damjan, Jelaska Sven D., Symondson William O. C. Prey detection in carabid beetles (Coleoptera: Carabidae) in woodland ecosystems by PCR analysis of gut contents. European Journal of Entomology. 2014;111(5):631–638. doi: 10.14411/eje.2014.079. DOI

Šerić Jelaska Lucija, Symondson W. O.C. Predation on epigeic, endogeic and anecic earthworms by carabids active in spring and autumn. Periodicum Biologorum. 2016;118(3):281–289. doi: 10.18054/pb.2016.118.3.4709. DOI

Šerić Jelaska L., Anđelić Dmitrović B., Ivanković Tatalović L., Rešetnik L. Meditera3. University of Zagreb Faculty of Science; 2022. DOI

Šerić Jelaska Lucija, Ivanković Tatalović Lara, Kostanjšek Fran, Kos Tomislav. Ground beetle assemblages and distribution of functional traits in olive orchards and vineyards depend on the agricultural management practice. BioControl. 2022;67(3):275–286. doi: 10.1007/s10526-022-10133-x. DOI

Skejo Josip, Rebrina Fran, Szövényi Gergely, Puskás Gellért, Tvrtković Nikola. The first annotated checklist of Croatian crickets and grasshoppers (Orthoptera: Ensifera, Caelifera) Zootaxa. 2018;4533(1) doi: 10.11646/zootaxa.4533.1.1. PubMed DOI

Štamol V., Erőss Z. P., Kletečki E., Vuković M. Terrestrial snails (Mollusca: Gastropoda) of islands of Šolta, Drvenik veli and Drvenik mali (Croatia. Natura Croatica: Periodicum Musei Historiae Naturalis Croatici. 2017;26(1):45–64. doi: 10.20302/NC.2017.26.4. DOI

Suwa M., Darvas B. Family Anthomyiidae. Papp L., Darvas B., editors. Contributions to a manual of Palaearctic Diptera Volume 3 1998

Teofilova T. M. Pseudomaquises in SW Bulgaria as a habitat for the ground beetles (Coleoptera: Carabidae) Zoology and Ecology. 2020;30(1):27–36. doi: 10.35513/21658005.2020.1.4. DOI

Trautner J., Geigenmüller K. Tiger beetles, ground beetles. Illustrated key to the Cicindelidae and Carabidae of Europe. J.Margraf Publishing; Aichtal: 1987. 488.

Vigna Taglianti A. Fauna Europaea: Coleoptera, Carabidae. Fauna Europaea version 2.6. https://fauna-eu.org/ https://fauna-eu.org/

Vujčić-Karlo S., Brigić A., Šerić Jelaska L., Kokan B., Hrašovec B. Crveni popis ugroženih vrsta trčaka (Coleoptera, Carabidae) u Hrvatskoj. Državni zavod za zaštitu prirode; Zagreb, Croatia: 2007.

Welter-Schultes F. W. European non-marine molluscs, a guide for species identification. Planet Poster Editions; Goettingen, Germany: 2012. 757

Wiktor A. Endemism of slugs within the Balkan Peninsula and adjacent islands (Gastropoda: Pulmonata: Arionidae. Milacidae, Limacidae, Agriolimacidae). Genus. 1997;8(1):205–221.

Winnertz J. Beitrag zu einer Monographie der Sciarinen. Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft; Wien, Austria: 1867.

Zicha O., et al. BioLib.cz. https://www.biolib.cz/ https://www.biolib.cz/