Extant representatives of the mayfly family Ameletopsidae Edmunds, 1957 as well as other three small families (Nesameletidae Riek, 1973, Oniscigastridae Lameere, 1917 and Rallidentidae Penniket, 1966) traditionally have been classified within the paraphyletic superfamily Siphlonuroidea. Except for Rallidentidae, which are endemic to New Zealand, they have an amphinotic distribution. Ameletopsidae are present with two genera in South America, one genus in Australia, and one genus in New Zealand. The fossil record of Ameletopsidae is scarce. The Mesozoic monospecific genus Promirara Jell & Duncan, 1986 was described from a larva of the Early Cretaceous Koonwarra Fossil Bed in Australia. Also associated with Ameletopsidae is the Cenozoic genus Balticophlebia Demoulin, 1968, which was recorded from Eocene Baltic amber based on a female adult. While the systematic position of the Eocene Balticophlebia seems to be still unclear and in need of clarification, we are able to confirm the presence of Ameletopsidae in the Eocene of Europe by describing Nebesna sotnia gen. & sp. nov. based on a relatively well-preserved male imago from Baltic amber. The fossil record thus indeed supports an ancient Pangean history of this family.

Biology Centre of the Czech Academy of Sciences Institute of Entomology Branišovská 31 37005 České Budějovice Czech Republic

Department of Entomology State Museum of Natural History Stuttgart Rosenstein 1 70191 Stuttgart Stuttgart Germany

Faculty of Biology and Environmental Protection Department of Invertebrate Zoology and Hydrobiology University of Lodz Banacha 12 16 90237 Łódź Poland

Independent Author Talino Str 33 1 05200 Vilnius Lithuania

National Museum of Natural History National Academy of Sciences of Ukraine Bohdan Khmelnytsky 15 Kyiv 01030 Ukraine

State Museum of Natural History National Academy of Sciences of Ukraine Teatralna 18 Lviv 79008 Ukraine

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