Permopsocids are small acercarian insects with mouthparts specialized for sucking. They are closely related to Hemiptera and Thysanoptera. The earliest known representatives are from the Early Permian. Here evidence is presented that the Permopsocida occurred even earlier in Pennsylvanian (Moscovian) deposits in the Piesberg quarry near Osnabrück (Lower Saxony, Germany). This material is assigned to the Permian family Psocidiidae; Carbonopsocus mercuryi gen. et sp. nov., based on the wing venation diagnosed by the unique branching pattern of the main veins, the shape of the areola postica being longer than wide, the angular shape of the pterostigma, the ir crossvein directed proximally mid of pterostigma (apomorphy) and the vannus formed by the three veins of PCu, A1 and A2. The shape of the veins, with a Y-vein formed by the distal fusion of PCu with A1, could be a putative symplesiomorphy of the Psocodea with Permopsocida and Hemiptera. C. mercuryi gen. et sp. nov. is the first appearance date for Permopsocida and roots the Acercaria tree. In addition, another specimen of Dichentomum cf. arroyo (Psocidiidae) from Carrizo Arroyo is presented and figured, confirming the presence of the genus Dichentomum near the Carboniferous-Permian boundary and linking it to the Artinskian species from Elmo in Kansas, USA.

Department of Zoology Faculty of Science Charles University Viničná 7 CZ 128 00 Praha 2 Czech Republic

Institut de Systématique Évolution Biodiversité Muséum national d'Histoire naturelle CNRS Sorbonne Université EPHE Université des Antilles CP50 57 rue Cuvier 75005 Paris France

Institute of Entomology Biology Centre of the Czech Academy of Sciences Branišovská 31 CZ 370 05 České Budějovice Czech Republic

Museum am Schölerberg Klaus Strick Weg 10 49082 Osnabrück Germany

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