The first phylogenetic inference of Calopterini and Eurrhacini focused on Calocladon and related taxa was carried out. A data matrix composed of 46 species and 51 morphological characters was assembled and analyzed using parsimony and model-based approaches. Eurrhacini were recovered monophyletic. Furthermore, phylogenetic analyses highly supported the Calocladon clade including also Atlanticolycus, Cladocalon, and Gorhamium gen. nov. as its sister clade. Our trees consistently recovered monophyly of the new genus with two new species: Gorhamiumbidentatum sp. nov. (Panama, Baru Volcano) and G.unidentatum sp. nov. from the Pacific slopes of Ecuador. A revised key to the genera of Eurrhacini is given and illustrations of distinguishing characters are provided. Phylogenetic relationships of Eurrhacini and character evolution are discussed.

Departamento de Engenharia Ambiental Universidade Estadual do Centro Oeste Rua Professora Maria Roza Zanon de Almeida s n Engenheiro Gutierrez Irati PR CEP 84505 677 Brazil Universidade Estadual do Centro Oeste Irati Brazil

Department of Biology Faculty of Education Palacky University Purkrabska 2 CZ 77140 Olomouc Czech Republic Palacky University Olomouc Czech Republic

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