The predaceous beetle family Thanerocleridae is one of the smallest families of Cleroidea. It comprises only 36 extant species widespread on all continents. Three more species have been described from Cretaceous ambers of Myanmar and France. The fourth fossil representative of Thanerocleridae is described herein. Thanerosus antiquus gen. and sp. nov. is based on one fossil specimen preserved in an amber piece from Upper Cretaceous Kachin amber. The holotype was imaged using an X-ray micro-CT system to obtain high-quality 3D images. A phylogenetic analysis based on 33 morphological characters supports the placement of the new genus at the basal position in a tree of Thanerocleridae, in the vicinity of extant Zenodosus Wolcott and three extinct Mesozoic genera with which the new fossil shares open procoxal and mesocoxal cavities and transverse procoxae. We offer here a key to all extant and extinct genera in the family together with a complete list of all valid thaneroclerid taxa.

Departament de Dinàmica de la Terra i de l'Oceà Facultat de Ciències de la Terra Universitat de Barcelona 08028 Barcelona Spain

Department of Entomology Moravian Museum 62700 Brno Czech Republic

Institut de Recerca de la Biodiversitat Universitat de Barcelona 08028 Barcelona Spain

Institute of Geosciences Section Paleontology University of Bonn 53115 Bonn Germany

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