The family Axymyiidae includes four extant genera and nine species known from the Holarctic and Oriental regions, with only one species Mesaxymyiakerteszi (Duda, 1930) occurring in Europe. The genus Plesioaxymyia Sinclair, 2013 was first discovered in Alaska in 1962, but officially described only many years later. Up to now, the genus included one species Plesioaxymyiavespertina Sinclair, 2013, known from two female specimens from western North America. During the study of the Diptera fauna in Paanajarvi National Park (Northwest Russia), one female specimen of Plesioaxymyia, was found. It differs from the Nearctic P.vespertina by details of its female terminalia and is recognized as a new species, herein described as Plesioaxymyiaimprevista sp. nov. The biology and geographic distribution of Plesioaxymyia is briefly discussed. The phylogenetic position of the genus, along with the family Axymyiidae is analyzed in the light of new molecular data, including sequences of three mitochondrial (ribosomal 12S and 16S and protein-encoding COI) and three nuclear genes (ribosomal 18S and 28S, protein-encoding CAD) for 72 terminal taxa. Axymyiidae is recovered as a monophyletic group with closest relatives in the infraorder Culicomorpha and Plesioaxymyiaimprevista sp. nov. representing the sister taxon to all the other species of Axymyiidae included in the analysis.

• Klíčová slova
• Biology, Nematocera, Russia, distribution, lower Diptera, new species, systematics,
• Publikační typ
• časopisecké články MeSH

Megagrapha starki Barták & Grootaert, sp. nov. (Poland, Russia, Slovakia), Oedalea portugalica Barták & Grootaert, sp. nov. (Portugal), Hybos conicus Grootaert & Barták, sp. nov. (Greece, Turkey), and Platypalpus obscuroides Barták & Grootaert, sp. nov. (Slovakia) are described and illustrated. Diagnostic characters are discussed. The female of Syndyas merzi Shamshev & Grootaert, 2012 is described for the first time. New distributional records are presented: Megagrapha europaea Papp & Földvári, 2001 is first reported from Slovakia and Syndyas merzi Shamshev & Grootaert, 2012 is first reported from Turkey.

• Klíčová slova
• Hybos, Megagrapha, Oedalea, Platypalpus, Syndyas, Descriptions, distribution, new species, re-description, taxonomy,
• Publikační typ
• časopisecké články MeSH

Atelestus turcicus Barták, sp. nov. (Turkey) and Atelestus ibericus Barták, sp. nov. (Spain) are described and illustrated. A key to all known Palaearctic species of Atelestus is provided and the main diagnostic characters are discussed. The female of Nemedina acutiformis Carles-Tolrá, 2008 is described for the first time. New distributional records are presented: Atelestus dissonans Collin, 1961 - first records from Spain and Bulgaria, A. pulicarius (Fallén, 1816) - first record from Turkey, Nemedina alamirabilis Chandler, 1981 - first record from Bulgaria and N. acutiformis Carles-Tolrá, 2008 - first record from Turkey.

• Klíčová slova
• Atelestidae, Atelestus, Diptera, Nemedina, Europe, descriptions, taxonomy,
• Publikační typ
• časopisecké články MeSH

The family Dolichopodidae forms two of the four largest evolutionary radiations in the Hawaiian Islands across all flies: Campsicnemus (183 spp) and the Eurynogaster complex (66 spp). They also include a small radiation of Conchopus (6 spp). A handful of other dolichopodid species are native to the islands in singleton lineages or small radiations. This study provides a phylogenetic perspective on the colonization history of the dolichopodid fauna in the islands. We generated a multi-gene data set including representatives from 11 of the 14 endemic Hawaiian dolichopodid genera to examine the history of colonization to the islands, and analyzed it using Bayesian and maximum likelihood phylogenetic methods. We used a subset of the data that included Conchopus and the eight genera comprising the Eurynogaster complex to estimate the first phylogenetic hypothesis for these endemic groups, then used Beast to estimate their age of arrival to the archipelago. The Eurynogaster complex, Campsicnemus and Conchopus are clearly the result of independent colonizations. The results strongly support the Eurynogaster complex as a monophyletic group, and also supports the monophyly of 4 of the 8 described genera within the complex (Adachia, Arciellia, Uropachys and Eurynogaster). Members of the family Dolichopodidae have been dispersing over vast distances to colonize the Hawaiian Archipelago for millions of years, leading to multiple independent evolutionary diversification events. The Eurynogaster complex arrived in the Hawaiian Archipelago 11.8 Ma, well before the arrival of Campsicnemus (4.5 Ma), and the even more recent Conchopus (1.8 Ma). Data presented here demonstrate that the Hawaiian Dolichopodidae both disperse and diversify easily, a rare combination that lays the groundwork for field studies on the reproductive isolating mechanisms and ecological partitioning of this group.

• Klíčová slova
• Colonization history, Diptera, Divergence dating, Dolichopodidae, Evolutionary radiation, Hawaiian islands, Long distance dispersal,
• Publikační typ
• časopisecké články MeSH

The phylogeny of the insect infraorder Bibionomorpha (Diptera) is reconstructed based on the combined analysis of three nuclear (18S, 28S, CAD) and three mitochondrial (12S, 16S, COI) gene markers. All the analyses strongly support the monophyly of Bibionomorpha in both the narrow (sensu stricto) and the broader (sensu lato) concepts. The major lineages of Bibionomorpha sensu lato (Sciaroidea, Bibionoidea, Anisopodoidea, and Scatopsoidea) and most of the included families are supported as monophyletic groups. Axymyiidae was not found to be part of Bibionomorpha nor was it found to be its sister group. Bibionidae was paraphyletic with respect to Hesperinidae and Keroplatidae was paraphyletic with respect to Lygistorrhinidae. The included Sciaroidea incertae sedis (except Ohakunea Edwards) were found to belong to one clade, but the relationships within this group and its position within Sciaroidea require further study.

• Klíčová slova
• Lower Diptera, Molecular markers, Phylogenetic analysis, Sciaroidea, Systematics,
• Publikační typ
• časopisecké články MeSH