Plant traits-the morphological, anatomical, physiological, biochemical and phenological characteristics of plants-determine how plants respond to environmental factors, affect other trophic levels, and influence ecosystem properties and their benefits and detriments to people. Plant trait data thus represent the basis for a vast area of research spanning from evolutionary biology, community and functional ecology, to biodiversity conservation, ecosystem and landscape management, restoration, biogeography and earth system modelling. Since its foundation in 2007, the TRY database of plant traits has grown continuously. It now provides unprecedented data coverage under an open access data policy and is the main plant trait database used by the research community worldwide. Increasingly, the TRY database also supports new frontiers of trait-based plant research, including the identification of data gaps and the subsequent mobilization or measurement of new data. To support this development, in this article we evaluate the extent of the trait data compiled in TRY and analyse emerging patterns of data coverage and representativeness. Best species coverage is achieved for categorical traits-almost complete coverage for 'plant growth form'. However, most traits relevant for ecology and vegetation modelling are characterized by continuous intraspecific variation and trait-environmental relationships. These traits have to be measured on individual plants in their respective environment. Despite unprecedented data coverage, we observe a humbling lack of completeness and representativeness of these continuous traits in many aspects. We, therefore, conclude that reducing data gaps and biases in the TRY database remains a key challenge and requires a coordinated approach to data mobilization and trait measurements. This can only be achieved in collaboration with other initiatives.

AgroParisTech Paris France

Aix Marseille Univ Univ Avignon CNRS IRD IMBE Marseille France

Algoma University Sault Ste Marie ON Canada

AMAP CIRAD CNRS IRD INRA Université de Montpellier Montpellier France

AMAP CIRAD CNRS IRD INRA Université de Montpellier Montpellier France AMAP IRD Herbier de Nouvelle Calédonie Nouméa New Caledonia

AMAP CIRAD CNRS IRD INRA Université de Montpellier Montpellier France French Institute of Pondicherry Puducherry India CEFE CNRS EPHE Université de Montpellier Université Paul Valéry Montpellier France

Applied Ecology Research Group School of Life Sciences Anglia Ruskin University Cambridge UK Faculty of Environmental Sciences and Natural Resource Management Norwegian University of Life Sciences Ås Norway

ARC Centre for Excellence in Plant Energy Biology Australian National University Acton ACT Australia

Archbold Biological Station Venus FL USA

Archbold Biological Station's Buck Island Ranch FL Lake Placid USA

Arizona State University Tempe AZ USA

Arnold Arboretum of Harvard University Boston MA USA

Ashiu Forest Research Station Field Science Education and Research Center Kyoto University Kyoto Japan

Barcelona School of Agricultural Engineering Universitat Politècnica de Catalunya Catalonia Spain

BayCEER Department of Disturbance Ecology University of Bayreuth Bayreuth Germany

BC Ministry Forest Lands Natural Resource Operations and Rural Development Dawson Creek BC Canada

Berlin Brandenburg Institute of Advanced Biodiversity Research Berlin Germany Freie Universität Berlin Berlin Germany

Biodiversity Conservation Laboratory Department of Environment University of the Aegean Mytilene Greece

Biodiversity Conservation Laboratory Department of Environment University of the Aegean Mytilene Greece Institute of Ecology and Evolution University of Bern Bern Switzerland

Biodiversity Macroecology and Biogeography University of Goettingen Göttingen Germany

Biodiversity Macroecology and Biogeography University of Goettingen Göttingen Germany Centre of Biodiversity and Sustainable Land Use University of Goettingen Göttingen Germany

Biological Sciences George Washington University Washington DC USA

Biological Sciences University of Tasmania Hobart Australia

Biology Department Oklahoma State University Oklahoma OK USA

Biology Department Wilfrid Laurier University Waterloo ON Canada

Biotechnical Faculty University of Ljubljana Ljubljana Slovenia

Bioversity International Lima Peru

Botanical Garden of the Samara University Samara Russia

Botany Department Faculty of Science Suez Canal University Ismailia Egypt

Botany Department University of Wyoming Laramie WY USA

Campus da Zapateira University of A Coruña A Coruña Spain

Carl von Ossietzky University of Oldenburg Oldenburg Germany

Carnegie Museum of Natural History Pittsburgh PA USA

CATIE Centro Agronómico Tropical de Investigación y Enseñanza Turrialba Costa Rica

CAVElab Computational and Applied Vegetation Ecology Ghent University Ghent Belgium

Center for Biodiversity Management Yungaburra Qld Australia

Center for Ecosystem Science and Society Northern Arizona University Flagstaff AZ USA

Center For Tree Science The Morton Arboretum Lisle IL USA

Centre d'Ecologie et des Sciences de la Conservation Muséum National d'Histoire Naturelle Centre National de la Recherche Scientifique Sorbonne Université Paris France

Centre for Applied Ecology Professor Baeta Neves School of Agriculture University of Lisbon Lisbon Portugal

Centre for Crop Systems Analysis Wageningen University Wageningen The Netherlands

Centre for Ecological Research and Forestry Applications Tervuren Belgium

Centre for Ecology and Hydrology Lancaster Environment Centre Lancaster UK

Centre for Ecology and Hydrology Wallingford Oxfordshire UK

Centre for Forest Research Institute for Integrative Systems Biology Université Laval Quebec QC Canada

Centre for Functional Ecology Departamento de Ciências da Vida Universidade de Coimbra Coimbra Portugal

Centre for Rainforest Studies The School for Field Studies Yungaburra Qld Australia

Centre of Excellence for Bioscurity Risk Analysis The University of Melbourne Melbourne Vic Australia

Centro de Investigación en Biodiversidad y Conservación Universidad Autónoma del Estado de Morelos Morelos Mexico

Centro de Modelación y Monitoreo de Ecosistemas Universidad Mayor Santiago Chile

Centro Politécnico Universidade Federal do Paraná Curitiba Brazil

Cibodas Botanical Garden Indonesian Institute of Sciences School Of Biosciences University of Melbourne Parkville Vic Australia

CIRAD UMR Eco and Sols Montpellier France Eco and Sols CIRAD INRA IRD SupAgro University of Montpellier Montpellier France

CIRAD UMR EcoFoG Kourou France

Cirad UMR EcoFoG Kourou French Guiana France

CIRAD UMR SELMET Montpellier France SELMET CIRAD INRA Univ Montpellier Montpellier SupAgro France

Cirad Université de Montpellier Montpellier France Institut National Polytehcnique Félix Houphouet Boigny INP HB Yamoussoukro Ivory Coast

Colgate University Hamilton NY USA

College Environmental Studies Yale University New Haven CT USA Department of Biological Sciences National University of Singapore Singapore Singapore Smithsonian Tropical Research Institute Panama City Panama

College of Chemistry and Life Sciences Zhejiang Normal University Jinhua China

College of Life and Environmental Sciences University of Exeter Penryn UK

College of Life Sciences Zhejiang University Hangzhou China

College of Resources and Environmental Sciences China Agricultural University Beijing China Xinjiang Institute of Ecology and Geography Chinese Academy of Sciences Urumqi China Research Center for Ecology and Environment of Central Asia Chinese Academy of Sciences Urumqi China

College of Science and Engineering James Cook University Smithfield Qld Australia

Conicet Consejo Nacional de Investigaciones Científicas y Técnicas Buenos Aires Argentina Facultad de Ciencias Forestales Universidad Nacional de Santiago del Estero Santiago del Estero Argentina

Conicet Consejo Nacional de Investigaciones Científicas y Técnicas Buenos Aires Argentina Facultad de Ingeniería y Ciencias Hídricas Universidad Nacional del Litoral Santa Fe Argentina

Consejo Nacional de Investigaciones Científicas y Técnicas Instituto Multidisciplinario de Biología Vegetal and Factulad de Ciencias Exactas Físicas y Naturales Universidad Nacional de Córdoba Córdoba Argentina

Conservation Ecology Groningen Institute for Evolutionary Life Sciences University of Groningen Groningen The Netherlands

CREA Research Centre for Forestry and Wood Arezzo Italy

CREAF Catalonia Spain Global Ecology Unit CREAF CSIC Universitat Autònoma de Barcelona Barcelona Spain

CREAF Catalonia Spain Universitat Autònoma de Barcelona Catalonia Spain

CSIC Institute for Sustainable Agriculture Cordoba Spain

CSIRO Canberra ACT Australia

CSIRO Land and Water Hobart Tas Tas Australia

Departamento de Biología Animal Biología Vegetal y Ecología Universidad de Jaén Jaén Spain

Departamento de Biología Universidad Nacional de Colombia Bogota Colombia

Departamento de Biología y Geología Física y Química Inorgánica y Analítica Universidad Rey Juan Carlos Móstoles Spain

Departamento de Botânica SCB UFPR Federal University of Parana Curitiba Brazil

Departamento de Botânica Universidade Federal de Pernambuco Recife PE Brazil

Departamento de Botânica Universidade Federal do Paraná Curitiba Brazil

Departamento de Ciencias Ambientales y Recursos Naturales Renovables Facultad de Ciencias Agronómicas Universidad de Chile Santiago Chile

Departamento de Ciencias de la Vida Universidad de las Fuerzas Armadas Sangolquí Ecuador

Departamento de Dinamica y Gestion Forestal INIA CIFOR Madrid Spain Sustainable Forest Management Research Institute University of Valladolid INIA Madrid Spain

Departamento de Ecología Centro de Investigación en Biodiversidad y Cambio Global Universidad Autónoma de Madrid Madrid Spain

Departamento de Ecología Funcional y Evolutiva Estación Experimental de Zonas Áridas La Cañada de San Urbano Spain

Departamento de Ecologia Universidade Federal do Rio Grande do Norte Natal RN Brazil

Département des Science Université du Québec À Montréal Montreal QC Canada

Department Computational Landscape Ecology UFZ Helmholtz Centre for Environmental Research Leipzig Germany Department Computational Hydrosystems UFZ Helmholtz Centre for Environmental Research Leipzig Germany Department of Landscape Architecture and Rural Systems Engineering Seoul National University Seoul Republic of Korea

Department for Molecular Biology and Genetics Democritus University of Thrace Alexandroupolis Greece

Department for Tisza River Research MTA Centre for Ecological Research DRI Debrecen Hungary

Department Land Environment Agriculture and Forestry Università degli Studi di Padova Padua Italy

Department of Agricultural and Environmental Sciences University of Milan Milano Italy

Department of Agriculture and Forest Engineering University of Lleida Lleida Spain Joint Research Unit CTFC AGROTECNIO Solsona Spain

Department of Animal and Plant Sciences University of Sheffield Sheffield UK

Department of Biogeography University of Bayreuth Bayreuth Germany

Department of Biological and Environmental Sciences University of Gothenburg Gothenburg Sweden

Department of Biological Sciences East Stroudsburg University East Stroudsburg PA USA

Department of Biological Sciences Florida International University Miami FL USA Oak Ridge National Laboratory US Department of Energy Oak Ridge TN

Department of Biological Sciences Goethe Universität Frankfurt Frankfurt am Main Germany Department of Botany and Molecular Evolution Senckenberg Research Institute and Natural History Museum Frankfurt am Main Germany

Department of Biological Sciences Macquarie University Sydney NSW Australia

Department of Biological Sciences Macquarie University Sydney NSW Australia Plant Sciences Forschungszentrum Jülich GmbH Jülich Germany

Department of Biological Sciences University of Calgary Calgary AB Canada

Department of Biological Sciences University of Cape Town Cape Town South Africa

Department of Biological Sciences University of Cape Town Cape Town South Africa SAEON Fynbos Node Claremont South Africa

Department of Biological Sciences University of Tennessee Knoxville TN USA Rocky Mountain Biological Laboratory Crested Butte CO USA

Department of Biology Colgate University Hamilton NY USA

Department of Biology Colorado State University Fort Collins CO USA

Department of Biology Duke University Durham NC USA

Department of Biology Faculty of Science Ferdowsi University of Mashhad Mashhad Iran

Department of Biology Federal University of Lavras Lavras MG Brazil

Department of Biology FFCLRP USP Ribeirão Preto Brazil

Department of Biology Hacettepe University Ankara Turkey

Department of Biology Stanford University Stanford CA USA

Department of Biology University of Copenhagen Copenhagen Denmark

Department of Biology University of Copenhagen Copenhagen Denmark Dept Animal Biology Plant Biology and Ecology University of Jaén Jaén Spain

Department of Biology University of Copenhagen Frederiksberg C Denmark

Department of Biology University of Florida Gainesville FL USA

Department of Biology University of Garmian Kalar Iraq School of Biological Sciences and Center for Ecology Southern Illinois University Carbondale Carbondale IL USA

Department of Biology University of Konstanz Konstanz Germany Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation Taizhou University Taizhou China

Department of Biology University of Maryland College Park MD USA

Department of Biology University of North Florida Jacksonville FL USA

Department of Biology University of Pisa Pisa Italy

Department of Biology University of Regina Regina SK Canada

Department of Biology University of Waterloo Waterloo ON Canada

Department of Biotechnology and Life Sciences University of Insubria Varese Italy

Department of Botany and Biodiversity Research Centre University of British Columbia Vancouver BC Canada

Department of Botany and Biodiversity Research University of Vienna Vienna Austria

Department of Botany and Biodiversity Research University of Vienna Vienna Austria Department of Ecology and Evolution University of Lausanne Lausanne Switzerland

Department of Botany and Molecular Evolution Senckenberg Research Institute and Natural History Museum Frankfurt am Main Germany

Department of Botany and Molecular Evolution Senckenberg Research Institute and Natural History Museum Frankfurt am Main Germany Entomology 3 Senckenberg Research Institute and Natural History Museum Frankfurt am Main Germany

Department of Botany and Zoology Masaryk University Brno Czech Republic

Department of Botany and Zoology Masaryk University Brno Czech Republic Institute of Botany Czech Academy of Sciences Třeboň Czech Republic

Department of Botany Biosciences Institute Federal University of Rio Grande do Sul Porto Alegre Brazil

Department of Botany Faculty of Science and Technology University of Debrecen Debrecen Hungary

Department of Botany Goa University Goa India Department of Ecology and Environmental Sciences Pondicherry University Puducherry India

Department of Botany University of Debrecen Hungary

Department of Botany University of Hawai'i Honolulu HI USA

Department of Botany University of Wisconsin Madison Madison WI USA

Department of Chemistry and Pharmacy University of Sassari Sassari Italy

Department of Chemistry Life Sciences and Environmental Sustainability University of Parma Parma Italy

Department of Earth and Environmental Sciences KU Leuven Leuven Belgium

Department of Earth and Environmental Sciences University of Manchester Manchester UK

Department of Ecology and Evolution State University of New York at Stony Brook Stony Brook NY USA

Department of Ecology and Evolutionary Biology University of California Los Angeles CA USA

Department of Ecology and Evolutionary Biology University of Connecticut Storrs CT USA

Department of Ecology and General Biology Faculty of Ecology and Environmental Sciences Technical University in Zvolen Zvolen Slovakia Department of Ecosystem Biology Faculty of Science University of South Bohemia Ceske Budejovice Czech Republic

Department of Ecology and Plant Geography Faculty of Biology Moscow Lomonosov State University Moscow Russia

Department of Ecology Evolution and Behavior University of Minnesota Minneapolis MN USA

Department of Ecology Evolution and Behavior University of Minnesota St Paul MN USA

Department of Ecology French Institute of Pondicherry Pondicherry India

Department of Ecology School of Horticulture and Plant Protection Yangzhou University Yangzhou China State Key Laboratory of Urban and Regional Ecology Research Centre for Eco Environmental Sciences Chinese Academy of Sciences Beijing China

Department of Ecology Universidade Federal do Rio Grande do Sul Porto Alegre RS Brazil

Department of Ecology University of Innsbruck Innsbruck Austria

Department of Ecology University of Innsbruck Innsbruck Austria Eurac Research Institute for Alpine Environment Bozen Bolzano Italy

Department of Environment Forest and Nature Lab Ghent University Gontrode Melle Belgium

Department of Environmental and Life Sciences Biology Karlstad University Karlstad Sweden

Department of Environmental Biology Sapienza University of Rome Rome Italy

Department of Environmental Science Institute for Water and Wetland Research Radboud University Nijmegen The Netherlands

Department of Environmental Science Policy and Management University of California Berkeley Berkeley CA USA

Department of Environmental Systems Science ETH Zürich Zürich Switzerland Swiss Federal Institute for Forest Snow and Landscape Research WSL Birmensdorf Switzerland Department of Ecology and Evolutionary Biology University of California Los Angeles CA USA

Department of Evolution Ecology and Organismal Biology University of California Riverside Riverside CA USA

Department of Evolutionary Biology Ecology and Environmental Sciences Biodiversity Research Institute Universitat de Barcelona Barcelona Spain

Department of Fisheries Wildlife and Conservation Biology University of Minnesota St Paul MN USA

Department of Forest Resources University of Minnesota St Paul MN USA

Department of Forest Resources University of Minnesota St Paul MN USA Hawkesbury Institute for the Environment Western Sydney University Sydney NSW Australia

Department of Forest Sciences Transilvania University of Brasov Brasov Romania BC3 Basque Centre for Climate Change Scientific Campus of the University of the Basque Country Leioa Spain

Department of Forestry Mississippi State University Starkville MS USA

Department of Geography and Geology Kingston University Kingston upon Thames UK

Department of Geography Environment and Society University of Minnesota Minneapolis MN USA

Department of Geography Environment and Spatial Sciences Michigan State University East Lansing MI USA

Department of Geography Humboldt University of Berlin Berlin Germany Biodiversity Macroecology and Biogeography University of Goettingen Göttingen Germany

Department of Geography King's College London London UK

Department of Geography Planning and Recreation Northern Arizona University Flagstaff AZ USA

Department of Geography University of Zurich Zürich Switzerland

Department of Geosciences and Natural Resources Western Carolina University Cullowhee NC USA

Department of Green Chemistry and Technology Ghent University Gent Belgium Department of Environment Ghent University Gent Belgium

Department of Horticulture and Landscape Architecture Colorado State University Fort Collins CO USA

Department of Invasion Ecology Institute of Botany Czech Academy of Sciences Průhonice Czech Republic

Department of Invasion Ecology Institute of Botany Czech Academy of Sciences Průhonice Czech Republic Department of Ecology Faculty of Science Charles University Prague Czech Republic

Department of Landscape and Biodiversity Norwegian Institute of Bioeconomy Research Ås Norway

Department of Landscape Architecture Planning and Management Swedish University of Agricultural Sciences Alnarp Sweden Gothenburg Botanical Garden Gothenburg Sweden Gothenburg Global Biodiversity Centre Gothenburg Sweden

Department of Landscape Architecture University of Sheffield Sheffield UK

Department of Life and Environmental Sciences Botany Division University of Cagliari Cagliari Italy

Department of Life Sciences Imperial College London Silwood Park Ascot UK

Department of Life Sciences University of Alcalá Alcala de Henares Spain

Department of Organismic and Evolutionary Biology Harvard University Cambridge MA USA

Department of Physical and Environmental Sciences University of Toronto Scarborough Toronto ON Canada

Department of Physical Geography Stockholm University Stockholm Sweden Bolin Centre for Climate Research Stockholm Sweden

Department of Plant and Environmental Sciences Weizmann Institute of Science Rehovot Israel

Department of Plant and Soil Sciences University of Pretoria Pretoria South Africa

Department of Plant Biology and Ecology University of the Basque Country UPV EHU Bilbao Spain

Department of Plant Biology Institute of Biology University of Campinas Campinas Brazil

Department of Plant Sciences University of Cambridge Cambridge UK

Department of Plant Sciences University of Saskatchewan Saskatoon SK Canada

Department of Range Management Faculty of Natural Resources and Marine Sciences Tarbiat Modares University Noor Iran

Department of Scientific Services Gorongosa National Park Beira Sofala Province Mozambique

Department of Soil and Plant Sciences University of Delaware Newark DE USA

Department of Terrestrial Ecology Netherlands Institute of Ecology Wageningen The Netherlands

Department of Wildlife Fish and Environmental Studies Swedish University of Agricultural Sciences Umeå Sweden Centre for African Conservation Ecology Department of Zoology Nelson Mandela University Port Elizabeth South Africa

Department of Zoology School of Mathematical and Natural Science University of Venda Thohoyandou South Africa Department of Biological Sciences DST NRF Centre of Excellence Percy FitzPatrick Institute of African Ornithology University of Cape Town Rondebosch South Africa

Department of Zoology University of British Columbia Vancouver BC Canada

Department of Zoology University of Oxford Oxford UK Balliol College University of Oxford Oxford UK

Department of Zoology University of Oxford Oxford UK Rocky Mountain Biological Laboratory Crested Butte CO USA

Desertification Research Center Valencia Spain

Dipartimento di Bioscienze Università degli Studi di Milano Milano Italy

Earth System Science Department Stanford University Stanford CA USA

Ecological Sciences Vrije Universiteit Amsterdam Amsterdam The Netherlands

Ecologie Systématique Evolution CNRS AgroParisTech University of Paris Sud Université Paris Saclay Orsay France

Ecology and Conservation Biology Institute of Plant Sciences University of Regensburg Regensburg Germany

Embrapa Amazônia Oriental Belém Brazil

Embrapa Recursos Genéticos e Biotecnologia Brasilia DF Brazil

Environmental and Climate Sciences Department Brookhaven National Laboratory NY Upton USA

Environmental Change Institute University of Oxford Oxford UK

Environmental Sciences Copernicus Institute of Sustainable Development Utrecht University Utrecht The Netherlands Centre for African Conservation Ecology Nelson Mandela University Port Elizabeth South Africa

Environmental Sciences Division and Climate Change Science Institute Oak Ridge National Laboratory Oak Ridge TN USA

Escuela de Biología Universidad de Costa Rica San José Costa Rica

Estación Experimental del Zaidín Consejo Superior de Investigaciones Científicas Granada Spain Estación Biológica de Doñana Consejo Superior de Investigaciones Científicas Sevilla Spain

Estonian University of Life Sciences Tartu Estonia

ETH Zurich Zurich Switzerland

Eurac Research Institute for Alpine Environment Bozen Bolzano Italy

Evolution and Ecology Research Centre and School of Biological Earth and Environmental Sciences UNSW Sydney Sydney NSW Australia

Evolutionary Ecology of Plants Department Biology Philipps University Marburg Marburg Germany Department of Bioscience University Salzburg Salzburg Austria

Facultad de Ciencias Forestales Universidad Nacional de Santiago del Estero Santiago del Estero Argentina

Facultad de Ciencias Naturales y Matemáticas Universidad del Rosario Bogota Colombia

Facultad de Ingeniería Agroindustrial Universidad de las Américas Quito Equador

Faculté de foresterie de géographie et de géomatique Université Laval Quebec QC Canada

Faculty of Agricultural and Environmental Sciences University of Rostock Rostock Germany

Faculty of Arts and Sciences Molecular Biology and Genetics Ordu University Ordu Turkey

Faculty of Biosciences and Aquaculture NORD University Steinkjer Norway Department of Biology Norwegian University of Science and Technology NTNU Trondheim Norway

Faculty of Environmental Sciences University of Life Sciences Prague Praha Suchdol Czech Republic Institute of Botany Czech Academy of Sciences Třeboň Czech Republic

Faculty of Forestry and Wood Sciences Czech University of Life Sciences Prague Czech Republic

Faculty of Sciences Charles University Praha Czech Republic

Federal University of Alagoas Maceió AL Brazil

Federal University of Pernambuco Recife PE Brazil

Flathead Lake Biological Station University of Montana Polson MT USA School of Sustainability Arizona State University Tempe AZ USA

Florida Museum of Natural History University of Florida Gainesville FL USA

Forest Ecology and Forest Management Group Wageningen University and Research Wageningen The Netherlands World Agroforestry Nairobi Kenya

Forest Ecology and Forest Management Group Wageningen University The Netherlands

Forest Rangeland and Fire Sciences Department University of Idaho Moscow ID USA

Forest Research Group INDEHESA University of Extremadura Plasencia Spain

Forest Sciences Centre Faculty of Forestry and Conservation Science University of British Columbia Vancouver BC Canada

Forest Sciences University of Göttingen Göttingen Germany Centre for Biodiversity and Sustainable Land use University of Göttingen Göttingen Germany

Forestry and Forest Products Research Institute Tsukuba Japan

Forestry College Beijing Forestry University Beijing China

Four Peaks Environmental Science and Data Solutions Wenatchee WA USA

Free University of Bozen Bolzano Bolzano Italy

Friedrich Schiller Universität Jena Jena Germany

Fundação Zoobotânica do Rio Grande do Sul Porto Allegre Brazil

Fundación Natura Bogota Colombia Departamento de Geociencias y Medio Ambiente Universidad Nacional de Colombia Medellin Colombia

Galápagos Verde 2050 Charles Darwin Foundation Charles Darwin Research Station Galapagos Ecuador

Gembloux Agro Bio Tech Biodiversity and landscape University of Liège Belgium

Geobotany Faculty of Biology University of Freiburg Freiburg im Breisgau Germany

Géopole de l'Université de Sherbrooke Quebec QC Canada

German Center for Integrative Biodiversity Research Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Geobotany and Botanical Garden Martin Luther University Halle Wittenberg Halle Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Helmholtz Centre for Environmental Research UFZ Halle Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Helmholtz Centre for Environmental Research UFZ Halle Germany Martin Luther University Halle Wittenberg Halle Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Helmholtz Centre for Environmental Research UFZ Leipzig Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Institute for Ecosystem Research Geobotany Kiel University Kiel Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Institute of Biology Geobotany and Botanical Garden Martin Luther University Halle Wittenberg Halle Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Institute of Ecology and Evolution Friedrich Schiller University Jena Jena Germany

German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany Smithsonian Tropical Research Institute Balboa Ancon Republic of Panama

German Center for Integrative Biodiversity Research Leipzig Germany

Global Academy of Agriculture and Food Security University of Edinburgh Midlothian Scotland

Global Change Research Institute AS CR Brno Czech Republic

Global Ecology Unit CREAF CSIC Universitat Autònoma de Barcelona Barcelona Spain

GLORIA Coordination Institute for Interdisciplinary Mountain Research Austrian Academy of Sciences and Department of Integrative Biology and Biodiversity Research University of Natural Resources and Life Sciences Vienna Vienna Austria

Graduate School of Agriculture Kyoto University Kyoto Japan Graduate School of Global Environmental Studies Kyoto University Kyoto Japan

Graduate School of Agriculture Kyoto University Kyoto Japan University of Florida Gainesville FL USA School of Molecular and Life Sciences Curtin University Perth WA Australia

Graduate School of Environment and Information Sciences Yokohama National University Yokohama Japan

Graduate School of Life and Environmental Sciences University of Tsukuba Tsukuba Japan

Graduate School of Life Sciences Tohoku University Sendai Japan

Great Lakes Forestry Centre Canadian Forest Service Natural Resources Canada Sault Ste Marie ON Canada

Groningen Institute of Archaeology University of Groningen Groningen The Netherlands

Grupo de Investigación en Biodiversidad y Dinámica de Ecosistémas Tropicales Universidad del Tolima Ibagué Colombia

Grupo de Investigación en Biodiversidad y Dinámica de Ecosistémas Tropicales Universidad del Tolima Ibagué Colombia Centro Forestal Tropical Bajo Calima Universidad del Tolima Buenaventura Colombia

Harry Butler Institute Murdoch University Perth WA Australia Dept of Geography and Environmental Studies Stellenbosch University Matieland Stellenbosch South Africa

Hawkesbury Institute for the Environment Western Sydney University Sydney NSW Australia

Hawkesbury Institute for the Environment Western Sydney University Sydney NSW Australia Plant Physiology Division Coconut Research Institute of Sri Lanka Lunuwila Sri Lanka

Health and Environmental Sciences Xi'an Jiaotong Liverpool University Suzhou Jiangsu China

Helmholtz Centre for Environmental Research UFZ Leipzig Germany

Herbarium Haussknecht Friedrich Schiller Universität Jena Jena Germany

Herencia Santa Cruz Bolivia

Hiroshima University Higashi Hiroshima Japan

Hokkaido University Sapporo Japan

Holden Arboretum Kirtland Ohio USA

Humboldt State University Arcata CA USA

ICREA Barcelona Spain CREAF Barcelona Spain

Illinois Natural History Survey Prairie Research Institute University of Illinois Champaign IL USA

Iluka Resources Perth WA Australia School of Biological Sciences The University of Western Australia Perth WA Australia

Imperial College London UK

INIA CIFOR Madrid Spain

INRA UEFP Cestas France

INRA Université Clermont Auvergne UMR PIAF Clermont Ferrand France

Institut d Estudis Andorrans Andorra

Institut de recherche en biologie végétale Montréal QC Canada Université de Montréal Montréal QC Canada

Institut de recherche sur les forêts Université du Québec en Abitibi Témiscamingue Rouyn Noranda QC Canada

Institut für Biologie Freie Universität Berlin Berlin Germany Berlin Brandenburg Institute of Advanced Biodiversity Research Berlin Germany

Institut für Physische Geographie Biogeography and Biodiversity Lab Goethe Universität Frankfurt Frankfurt am Main Germany

Institut National Polytechnique Félix Houphouët Boigny Yamoussoukro Côte d'Ivoire

Institute Botanic Garden Ural Branch of the Russian Academy of Sciences Ekaterinburg Russia Tyumen State University Tyumen Russia

Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam The Netherlands

Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam The Netherlands Forest Ecology and Forest Management Group Wageningen University and Research Wageningen The Netherlands Institute for Global Ecology Florida Institute of Technology Melbourne FL USA

Institute for Ecosystem Research Geobotany Kiel University Kiel Germany

Institute for Plant Ecology Justus Liebig University Giessen Germany

Institute for Resources Environment and Sustainability University of British Columbia Vancouver BC Canada

Institute of Applied Ecology University of Canberra Canberra ACT Australia

Institute of Arctic Biology University of Alaska Fairbanks Fairbanks AK USA

Institute of Atmospheric and Earth System Research University of Helsinki Helsinki Finland

Institute of Biodiversity and Ecosystem Research Bulgarian Academy of Sciences Sofia Bulgaria

Institute of Biology and Environmental Sciences University of Oldenburg Oldenburg Germany Department of Biology Vrije Universiteit Brussel Brussels Belgium

Institute of Biology Bucharest Romanian Academy Bucharest Romania Institute of Ecology and Botany MTA Centre for Ecological Research Vácrátót Hungary

Institute of Biology Geobotany and Botanical Garden Martin Luther University Halle Wittenberg Halle Germany

Institute of Biology of Komi Science Centre of the Ural Branch of the Russian Academy of Sciences Syktyvkar Komi Republic Russia

Institute of Botany Czech Academy of Sciences Třeboň Czech Republic

Institute of Botany Czech Academy of Sciences Třeboň Czech Republic Department of Botany Faculty of Sciences University of South Bohemia Ceske Budejovice Czech Republic

Institute of Botany Czech Academy of Sciences Třeboň Czech Republic Faculty of Sciences Charles University Praha Czech Republic

Institute of Botany Plant Science and Biodiversity Center Slovak Academy of Sciences Bratislava Slovakia

Institute of Botany The Czech Academy of Sciences Prùhonice Czech Republic Department of Botany Faculty of Science Charles University Prague Czech Republic

Institute of Botany University of Natural Resources and Life Sciences Vienna Austria

Institute of Dendrology Polish Academy of Sciences Kornik Poland Department of Game Management and Forest Protection Faculty of Forestry Poznan University of Life Sciences Poznan Poland

Institute of Desertification Studies Chinese Academy of Forestry Beijing China

Institute of Ecology and Earth Sciences University of Tartu Tartu Estonia

Institute of Ecology and Evolution Friedrich Schiller University Jena Jena Germany

Institute of Ecology Tallinn University Tallinn Estonia

Institute of Environmental Sciences Leiden University Leiden The Netherlands

Institute of Evolutionary Ecology and Conservation Genomics Ulm University Ulm Germany

Institute of Geography and Geoecology Karlsruhe Institute of Technology Karlsruhe Germany

Institute of Geography Fujian Normal University Fuzhou China

Institute of Geography University of Erlangen Nuremberg Erlangen Germany

Institute of Integrative Biology ETH Zürich Zürich Switzerland

Institute of Landscape and Plant Ecology University of Hohenheim Stuttgart Germany

Institute of Lowland Forestry and Environment University of Novi Sad Novi Sad Serbia

Institute of Plant Sciences Bern Switzerland

Institute of Systematic Botany and Ecology Ulm University Ulm Germany

Institute of Temperate Forest Sciences Ripon QC Canada UQO Department of Natural Sciences Ripon QC Canada

Instituto Alexander Von Humboldt Bogota Colombia

Instituto de Biociências Laboratory of Vegetation Ecology Universidade Estadual Paulista Rio Claro Brazil

Instituto de Biología Tercer Circuito s n de Ciudad Universitaria Mexico City Mexico Universidad Nacional Autónoma de México Coyoacán Mexico

Instituto de Ciencias Ambientales y Evolutivas Universidad Austral de Chile Valdivia Chile

Instituto de Ciencias Naturales Universidad Nacional de Colombia Bogota Colombia Fundación Natura Bogota Colombia

Instituto de Investigación de Recursos Biológicos Alexander von Humboldt Bogota Colombia Universidad Distrital Francisco José de Caldas Bogota Colombia

Instituto de Investigaciones en Biodiversidad y Medioambiente CONICET Bariloche Argentina Departamento de Ecología Universidad Nacional del Comahue Bariloche Argentina

Instituto de Investigaciones en Biodiversidad y Medioambiente CONICET Universidad Nacional del Comahue Bariloche Argentina

Instituto de Investigaciones en Recursos Naturales Agroecología y Desarrollo Rural Universidad Nacional de Río Negro El Bolsón Argentina Conicet Consejo Nacional de Investigaciones Científicas y Técnicas Buenos Aires Argentina

Instituto de Recursos Naturales y Agrobiología de Salamanca Salamanca Spain

Instituto Federal de Educação Ciência e Tecnologia do Ceará Crateús Brazil

Instituto Nacional de Tecnología Agropecuaria CONICET Rio Gallegos Argentina

Instituto Nacional de Tecnología Agropecuaria Consejo Nacional de Invetigaciones Científicas y Técnicas Universidad Nacional de La Patagonia Austral Río Gallegos Argentina

Instituto Pirenaico de Ecología Zaragoza Spain

IRSTEA Aix en Provence UMR RECOVER Aix Marseille University Aix en Provence France Department of Environmental Systems Science ETH Zürich Zürich Switzerland Swiss Federal Institute for Forest Snow and Landscape Research WSL Birmensdorf Switzerland

Isotope Bioscience Laboratory ISOFYS Ghent University Gent Belgium

Jomo Kenyatta University of Agriculture and Technology Nairobi Kenya

Jonah Ventures Boulder CO USA

Julius von Sachs Institute for Biological Sciences Chair of Ecophysiology and Vegetation Ecology University of Wuerzburg Wuerzburg Germany

Jungwon University Goesan Chungbuk Korea

Kenyon College Gambier OH USA

Kochi University Nankoku Japan

Komarov Botanical Institute RAS St Petersburg Russia

Kyoto University Kyoto Japan

Laboratoire d'Ecologie Alpine Université Grenoble Alpes Grenoble Cedex 9 France French Institute of Pondicherry Puducherry India

Laboratoire ECODIV URA IRSTEA EA 1293 Normandie Université UFR ST Université de Rouen Mont Saint Aignan France

Laboratório de Ecologia Funcional de Comunidades Departamento de Botânica Universidade Federal do Paraná Curitiba Brazil

Laboratório de Ecologia Vegetal Departamento de Ecologia Programa de Pós Graduação em Ecologia Universidade Federal do Rio Grande do Sul Porto Alegre RS Brazil

Laboratório de Ecologia Vegetal Porto Alegre RS Brazil

Laboratório de Ecologia Vegetal Programa de Pós Graduação em Ecologia Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

Laboratório de Estudos em Vegetação Campestre Programa de Pós Graduação em Botânica Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

Laboratorio Nacional de Ciencias de la Sostenibilidad Instituto de Ecología Universidad Nacional Autónoma de México Ciudad Universitaria Mexico City Mexico

Lakehead University Thunder Bay ON Canada

Lancaster Environment Centre Lancaster University Lancaster UK

Landscape Ecology Group Institute of Biology and Environmental Sciences University of Oldenburg Oldenburg Germany

Macquarie University North Ryde NSW Australia

Manaaki Whenua Landcare Research Hamilton New Zealand

Manaaki Whenua Landcare Research Lincoln New Zealand

Max Planck Institute for Biogeochemistry Jena Germany

Max Planck Institute for Biogeochemistry Jena Germany German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany

Max Planck Institute for Biogeochemistry Jena Germany German Center for Integrative Biodiversity Research Halle Jena Leipzig Leipzig Germany University of Leipzig Leipzig Germany

McGill University Montreal QC Canada

Meijo University Nagoya Aichi Japan

Ministry of Education Key Laboratory for Earth System Modeling Department of Earth System Science Tsinghua University Beijing China

Morton Arboretum Lisle IL USA

MTA Centre for Ecological Research Tihany Hungary

MTA DE Lendület Functional and Restoration Ecology Research Group Debrecen Hungary Department of Ecology University of Debrecen Debrecen Hungary

MTA TKI Biodiversity and Ecosystem Services Research Group Debrecen Hungary

Museo Nacional de Ciencias Naturales CSIC Madrid Spain

Museu Paraense Emilio Goeldi Belém PA Brazil

National Institute for Research Development in Forestry Voluntari Romania

National Institute of Amazonian Research Manaus Brazil

National Taiwan University Taipei Taiwan

Natural Resources Canada Canadian Forest Service Northern Forestry Centre Edmonton AB Canada

Natural Resources Canada Canadian Forest Service Québec QC Canada

Natural Resources Canada Canadian Wood Fibre Centre Quebec QC Canada

Natural Resources Institute Finland Espoo Finland

Naturalis Biodiversity Center Leiden The Netherlands

Nelson Mandela University Port Elizabeth South Africa

New Jersey Institute of Technology Newark NJ USA

Norwegian Institute for Nature Research Trondheim Norway

Norwegian University of Science and Technology NTNU Trondheim Norway

NSW Department of Primary Industries Parramatta NSW Australia

Numerical Terradynamic Simulation Group College of Forestry and Conservation University of Montana Missoula USA

Oak Ridge National Laboratory Oak Ridge TN USA

Ontario Ministry of Natural Resources and Forestry Centre for Northern Forest Ecosystem Research Thunder Bay ON Canada

Osaka City University Osaka Japan

Osaka Natural History Center Osaka Japan

Oxford University Oxford UK

Pacific Northwest National Laboratory College Park MD USA

Pacific Northwest National Laboratory Joint Global Change Research Institute College Park MD USA

Pacific Northwest National Laboratory Richland WA USA

Pacific Northwest National Laboratory Richland WA USA University of Massachusetts Amherst Amherst MA USA

Peking University Beijing China

PERSYST Department CIRAD Montpellier Cedex 5 France

PIAF INRA Université Clermont Auvergne Clermont Ferrand France

Plant Conservation and Population Biology Department of Biology KU Leuven Leuven Belgium Division of Ecology Evolution and Biodiversity Conservation Heverlee Belgium

Plant Ecology University of Bayreuth Bayreuth Germany

Plant Ecophysiology and Evolution Group Guangxi Key Laboratory of Forest Ecology and Conservation College of Forestry Guangxi University Nanning Guangxi PR China State Key Laboratory for Conservation and Utilization of Subtropical Agro bioresources Guangxi University Nanning PR China

Pós graduação em Ecologia e Conservação Universidade Federal do Paraná Curitiba PR Brazil

Programa de pós graduação em Ecologia Universidade Federal do Rio Grande do Sul Porto Alegre RS Brasil

Programa Nacional de Pós Doutorado Programa de Pós Graduação em Ecologia Institute of Biology University of Campinas UNICAMP Brazil

Quantitative Plant Ecology and Biodiversity Research Laboratory Department of Biology Faculty of Science Ferdowsi University of Mashhad Mashhad Iran

Queensland University of Technology Brisbane Australia

Remote Sensing Laboratories Department of Geography University of Zurich Zurich Switzerland Research Unit Community Ecology Swiss Federal Institute for Forest Snow and Landscape Research WSL Birmensdorf Switzerland Department of Research and Geoinformation Swiss National Park Chastè Planta Wildenberg Zernez Switzerland

Research Institute for Humanity and Nature Kyoto Japan Center for Southeast Asian Studies Kyoto University Kyoto Japan

Research School of Biology The Australian National University Canberra ACT Australia School of Geosciences The University of Edinburgh Edinburgh UK

Research Unit ECOBIO Ecosystèmes Biodiversité Evolution Université Rennes 1 CNRS Rennes France

Restoration Ecology Technische Universität München München Germany

River Basin Research Center Gifu University Gifu Gifu Japan

Rocky Mountain Biological Laboratory Crested Butte CO USA Maritime and Science Technology Academy Miami FL USA

Rocky Mountain Biological Laboratory Crested Butte CO USA School of Life Sciences Arizona State University Tempe AZ USA

Royal Botanic Gardens Kew Richmond UK

Royal Botanic Gardens Kew West Sussex UK

Royal Museum for Central Africa Tervuren Belgium

Rubenstein School of Environment and Natural Resources University of Vermont Burlington VT USA

Rutgers University Piscataway NJ USA

SAEON Fynbos Node Claremont South Africa Department of Conservation Ecology and Entomology Stellenbosch University Matieland South Africa

Samara National Research University Samara Russia

Santa Catarina State University Lages SC Brazil

Sapienza University of Rome Rome Italy

School of Agriculture and Biology Shanghai Jiao Tong University Shanghai P R China Shanghai Urban Forest Ecosystem Research Station National Forestry and Grassland Administration Shanghai P R China

School of Animal Plant and Environmental Sciences University of the Witwatersrand Johannesburg South Africa

School of Biological Earth and Environmental Sciences UNSW Sydney Sydney NSW Australia

School of Biological Sciences and School of Agriculture and Environment The University of Western Australia Crawley WA Australia

School of Biological Sciences Southern Illinois University Carbondale Carbondale IL USA

School of Biological Sciences The University of Adelaide Adelaide SA Australia

School of Biological Sciences The University of Queensland Brisbane Qld Australia CSIRO Canberra ACT Australia Manaaki Whenua Landcare Research Lincoln New Zealand

School of Biological Sciences The University of Western Australia Crawley WA Australia

School of Biological Sciences University of Aberdeen Aberdeen UK

School of Biological Sciences University of Southampton Southampton UK Aquatic Ecology and Environmental Biology Radboud University Nijmegen Nijmegen The Netherlands

School of Biology Department of Botany Aristotle University of Thessaloniki Greece

School of Biosciences and Veterinary Medicine Plant Diversity and Ecosystems Management Unit University of Camerino Camerino Italy

School of Biosciences and Veterinary Medicine Plant Diversity and Ecosystems Management Unit University of Camerino Camerino Italy BIGEA Department of Biological Geological and Environmental Sciences Alma Mater Studiorum University of Bologna Bologna Italy

School of Civil and Environmental Engineering Yonsei University Seoul Korea

School of Earth and Environment Science University of Manchester Manchester UK

School of Environmental Sciences University of Guelph Guelph ON Canada

School of Forest Resources and Conservation University of Florida Gainesville FL USA

School of Forestry and Environmental Studies Yale University New Haven CT USA

School of Geography Earth and Environmental Sciences University of Birmingham Birmingham UK

School of Geography Geology and Environment University of Leicester Leicester UK

School of Geography University of Leeds Leeds UK

School of Geography University of Leeds Leeds UK Department of Geography University College London London UK

School of Geography University of Leeds Leeds UK School of Geography Earth and Environmental Sciences University of Birmingham Birmingham UK

School of Geography University of Nottingham Nottingham UK

School of Geosciences Edinburgh University Edinburgh UK

School of Geosciences The University of Edinburgh Edinburgh UK

School of Informatics Computing and Cyber Systems Northern Arizona University Flagstaff AZ USA

School of Life Science and Engineering Southwest University of Science and Technology Mianyang China Department of Crop and Forest Sciences and Agrotecnio Center Universitat de Lleida Lleida Spain

School of Life Sciences University of Sussex Brighton UK

School of Molecular and Life Sciences Curtin University Perth WA Australia College of Science Health Engineering and Education Murdoch University Murdoch WA Australia

School of Natural Sciences Bangor University Bangor UK

School of Plant Sciences and Food Security George S Wise Faculty of Life Sciences Tel Aviv University Tel Aviv Israel

Section for Ecoinformatics and Biodiversity Department of Bioscience Aarhus University Aarhus Denmark

Section for Ecoinformatics and Biodiversity Department of Bioscience Aarhus University Aarhus Denmark Center for Biodiversity Dynamics in a Changing World Department of Bioscience Aarhus University Aarhus Denmark

SELMET CIRAD INRA Univ Montpellier Montpellier SupAgro France CIRAD UMR SELMET PZZS Dakar Senegal

Senckenberg Biodiversität und Klima Forschungszentrum Frankfurt Germany Palmengarten der Stadt Frankfurt am Main Frankfurt Germany

Senckenberg Biodiversity and Climate Research Centre Frankfurt am Main Germany

Senckenberg Biodiversity and Climate Research Centre Frankfurt am Main Germany Department of Biological Sciences Goethe Universität Frankfurt Frankfurt am Main Germany

Senckenberg Biodiversity and Climate Research Centre Frankfurt am Main Germany Department of Physical Geography Goethe University Frankfurt am Main Germany

Sharplex Services Edgecomb ME USA

Smithsonian Conservation Biology Institute Front Royal VA USA

Smithsonian Environmental Research Center Edgewater MD USA

Smithsonian Tropical Research Institute Balboa Ancon Republic of Panama

Smithsonian Tropical Research Institute Balboa Ancon Republic of Panama Institute for Biology and Environmental Sciences University Oldenburg Oldenburg Germany

South African National Biodiversity Institute Pretoria South Africa

South China Botanical Garden Chinese Academy of Sciences Guangzhou China

Spanish National Research Council CSIC Catalonia Spain CREAF Catalonia Spain

State Key Laboratory of Grassland Agro ecosystems College of Pastoral Agriculture Science and Technology Lanzhou University Lanzhou China

State University of Campinas UNICAMP Campinas SP Brazil

Stockholm University Stockholm Sweden

Sun Yat sen University Guangzhou China

SUNY College of Environmental Science and Forestry Syracuse NY USA

Swedish Species Information Centre Swedish University of Agricultural Sciences Uppsala Sweden

Swedish Species Information Centre Swedish University of Agricultural Sciences Uppsala Sweden Department of Ecology Swedish University of Agricultural Sciences Uppsala Sweden

Swiss Federal Institute for Forest Snow and Landscape Research WSL Birmensdorf Switzerland

Swiss Federal Institute for Forest Snow and Landscape Research WSL Birmensdorf Switzerland Faculty of Forestry and Wood Sciences Czech University of Life Sciences Prague Czech Republic Institute of Agricultural Sciences Department of Environmental Systems Science ETH Zurich Zurich Switzerland

Systematic Botany and Functional Biodiversity Institute of Biology Leipzig University Leipzig Germany

Systems Ecology Department of Ecological Science Vrije Universiteit Amsterdam The Netherlands

Tartu Observatory University of Tartu Tartumaa Estonia

Technical University of Munich Freising Germany

Technische Universität Ilmenau Ilmenau Germany

Terrestrial Ecology Section Department of Biology University of Copenhagen Denmark

Teshio Experimental Forest Hokkaido University Horonobe Japan

Texas Tech University Lubbock TX USA

The James Hutton Institute Aberdeen UK

The James Hutton Institute Dundee UK King Saud University Riyadh Saudi Arabia

The Morton Arboretum Lisle IL USA

The Morton Arboretum Lisle IL USA The Field Museum Chicago IL USA

The University of Arizona Tucson AZ USA

The University of Manchester Manchester UK

The University of Western Australia Crawley WA Australia

Theoretical and Experimental Ecology Station CNRS Paul Sabatier University Toulouse Moulis France

Tulipan s n Móstoles Madrid Spain

TUM School of Life Sciences Weihenstephan Technical University of Munich Freising Germany

UCA INRA VetAgro Sup UMR Ecosystème Prairial Clermont Ferrand France

UCA INRA VetAgro Sup UMR Ecosystème Prairial Clermont Ferrand France Departamento de Biología y Geología Física y Química Inorgánica Escuela Superior de Ciencias Experimentales y Tecnología Universidad Rey Juan Carlos Móstoles Spain

UMR 5175 CEFE Univ Montpellier CNRS EPHE IRD Univ Paul Valéry Montpellier France

Unit for Modelling of Climate and Biogeochemical Cycles UR SPHERES University of Liège Liège Belgique

Unit of research SPHERES University of Liège Liège Belgium

Unit of Research SPHERES University of Liège Liège Belgium

Univ Bordeaux INRAE BIOGECO Cestas France

Univ Grenoble Alpes CNRS Univ Savoie Mont Blanc LECA Grenoble France

Universidad de Jaén Jaén Spain

Universidad Distrital Francisco José de Caldas Bogota Colombia Laboratorio de invasiones Biológicas Universidad de Concepcion Concepcion Chile

Universidad Nacional Autónoma de México Coyoacán Mexico

Universidade Federal de Pernambuco Recife PE Brazil

Universidade Federal do Acre Rio Branco AC Brazil

Universidade Federal do Paraná Curitiba PR Brazil

Universidade Federal do Rio Grande do Norte UFRN Natal RN Brazil

Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

Universidade Regional de Blumenau Blumenau SC Brazil

Universitat Autònoma de Barcelona Barcelona Spain

Universitat Politècnica de Catalunya Castelldefels Spain

Université de Lorraine AgroParisTech INRAE UMR Silva Nancy France

Université de Lorraine Lorraine France

Université de Sherbrooke Sherbrooke QC Canada

Université du Québec à Trois Rivières Trois Rivières Canada

Université Laval Quebec QC Canada

Université Libre de Bruxelles Bruxelles Belgium

University Centre Myerscough Preston UK

University of Bayreuth Bayreuth Germany

University of California at Berkeley Berkeley CA USA

University of California Irvine Irvine CA USA Southwest Biological Science Center U S Geological Survey Moab UT USA

University of California Riverside Riverside CA USA

University of California Santa Cruz Santa Cruz CA USA

University of Copenhagen Copenhagen Denmark

University of Florida Gainesville FL USA

University of Guelph Guelph ON Canada

University of Guelph Guelph ON Canada Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

University of Helsinki Helsinki Finland

University of Illinois at Chicago Chicago IL USA

University of Illinois at Urbana Champaign Urbana Champaign IL USA

University of Innsbruck Innsbruck Austria

University of Jaén Jaén Spain

University of Leipzig Leipzig Germany

University of Maine Orono ME USA

University of Maryland College Park MD USA

University of Minnesota St Paul MN USA

University of New England Armidale NSW Australia

University of New Hampshire Durham NH USA

University of Pittsburgh Pittsburgh PA USA

University of Reading Reading UK

University of Roma Tre Rome Italy

University of Science Vietnam National University Ho Chi Minh City Ho Chi Minh City Vietnam

University of Tartu Tartu Estonia

University of Tasmania Hobart Tas Australia

University of Toronto Scarborough Scarborough ON Canada

University of Toronto Toronto ON Canada

University of Vermont Burlington VT USA

University of Victoria Victoria BC Canada

University of Winnipeg Winnipeg Manitoba Canada

University of Wisconsin Eau Claire Eau Claire WI USA

University Pablo de Olavide Seville Spain

Ural State Forest Engineering University Ekaterinburg Russia Botanical Garden of Ural Branch of Russian Academy of Sciences Ekaterinburg Russia

USDA ARS Rangeland Resources and Systems Research Unit Fort Collins CO USA

Wageningen Environmental Research Wageningen The Netherlands

Wageningen University and Research Wageningen The Netherlands

Wageningen University and Research Wageningen The Netherlands Land Life Company Amsterdam The Netherlands

Water Management and Systems Research Unit United States Department of Agriculture Agricultural Research Service Fort Collins CO USA

WSL Swiss Federal Research Institute Birmensdorf Switzerland Faculty of Forestry University of British Columbia Vancouver BC Canada

Yamagata University Yamagata Japan

Yangzhou University Yangzhou Jiangsu China

Zoology Department Edward Grey Institute Oxford University Oxford UK Department of Zoology Cambridge University Cambridge Conservation Initiative Cambridge UK