The small, monophyletic mayfly family Vietnamellidae Allen, 1984 has so far only been known from a few extant species of the genus Vietnamella Tshernova, 1972, which are all distributed in the Oriental Realm (Vietnam, Thailand, China, and India). Herein we report the first fossil record of Vietnamellidae based on a male and female imago from Mid-Cretaceous Burmese amber. We establish the new genus Burmella gen. nov. to accommodate these two new Mesozoic specimens. Their attribution to Vietnamellidae is supported by the rounded shape of the hind wings with arched outer margin, the course of thoracic sutures, and characteristics of venation, especially of MP and Cu of the forewings and associated intercalary veins of the cubital field. At the same time, Burmella gen. nov. clearly differs from Vietnamella by a diminished number of longitudinal and cross veins in the hind wings, and by the different shape of male genitalia. This first fossil record of Vietnamellidae supports an age of at least 100 Ma for this taxon.

Biology Centre of the Czech Academy of Sciences Institute of Entomology Branišovská 31 37005 České Budějovice Czech Republic Biology Centre of the Czech Academy of Sciences Institute of Entomology České Budějovice Czech Republic

Department of Entomology State Museum of Natural History Stuttgart Rosenstein 1 70191 Stuttgart Germany State Museum of Natural History Stuttgart Stuttgart Germany

Department of Invertebrate Zoology and Hydrobiology University of Łódź Banacha 12 16 90237 Łódź Poland University of Łódź Łódź Poland

National Museum of Natural History National Academy of Sciences of Ukraine Bohdan Khmelnytsky 15 01030 Kyiv Ukraine National Museum of Natural History National Academy of Sciences of Ukraine Kyiv Ukraine

State Museum of Natural History National Academy of Sciences of Ukraine Teatralna 18 79008 Lviv Ukraine State Museum of Natural History National Academy of Sciences of Ukraine Lviv Ukraine

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