The monophyletic mayfly family Vietnamellidae has been introduced for a few extant species from the Indomalayan Realm. All these species belong to the genus Vietnamella and have been described in adult and larval stages. Recently, the fossil genus Burmella was established for male and female imagines of two new species from mid-Cretaceous Burmese amber and assigned to the family Vietnamellidae. In this contribution, we describe another species in the genus Burmella, namely B. inconspicua sp. nov. based on a female imago. It clearly differs from the previously described female of B. clypeata by the absence of an extension of the clypeus and by a different venation in fore and hind wings. Furthermore, we discuss here in detail the systematic position of the monotypic species Burmaheptagenia zhouchangfai originally assigned to the family Heptageniidae and provide arguments for the synonymy of the genus Burmaheptagenia syn. nov. with Burmella. We propose the new combination Burmella zhouchangfai comb. nov., together with a modified generic diagnosis for the genus Burmella and discuss the adult characters of Vietnamellidae. Systematic placement of the genus Burmella in Vietnamellidae and the phylogenetic position of this family within Ephemerelloidea are discussed. We also consider the geographic origin of the family and the role of the Cretaceous Burma Terrane migration as a key event that facilitated the transfer of Gondwanan fauna to Asia.

Biology Centre of the Czech Academy of Sciences Institute of Entomology České Budějovice Czech Republic

Department of Entomology State Museum of Natural History Stuttgart Stuttgart Germany

Department of Invertebrate Zoology and Hydrobiology Faculty of Biology and Environmental Protection University of Lodz Łódź Poland

Institute of Technology and Life Sciences National Research Institute Raszyn Poland

National Museum of Natural History National Academy of Sciences of Ukraine Kyiv Ukraine

School of Grain Science and Technology Jiangsu University of Science and Technology Zhenjiang China

State Museum of Natural History National Academy of Sciences of Ukraine Lviv Ukraine

The Key Laboratory of Jiangsu Biodiversity and Biotechnology College of Life Sciences Nanjing Normal University Nanjing China

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