Disorders of the brain can exhibit considerable epidemiological comorbidity and often share symptoms, provoking debate about their etiologic overlap. We quantified the genetic sharing of 25 brain disorders from genome-wide association studies of 265,218 patients and 784,643 control participants and assessed their relationship to 17 phenotypes from 1,191,588 individuals. Psychiatric disorders share common variant risk, whereas neurological disorders appear more distinct from one another and from the psychiatric disorders. We also identified significant sharing between disorders and a number of brain phenotypes, including cognitive measures. Further, we conducted simulations to explore how statistical power, diagnostic misclassification, and phenotypic heterogeneity affect genetic correlations. These results highlight the importance of common genetic variation as a risk factor for brain disorders and the value of heritability-based methods in understanding their etiology.

1st and 3rd Departments of Neurology Aristotle University of Thessaloniki Thessaloniki Greece

Aarhus Genome Center Aarhus Denmark

Academic Centre on Rare Diseases University College Dublin Dublin Ireland

ADHD Outpatient Clinic Hospital de Clínicas de Porto Alegre Porto Alegre Brazil

Advanced Interventions Service NHS Tayside Dundee UK

Aglaia Kyriakou Childrens Hospital Athens Greece

Alexandru Obregia Clinical Psychiatric Hospital Biometric Psychiatric Genetics Research Unit Bucharest Romania

Alpert Medical School of Brown University Providence RI USA

Altrecht Eating Disorders Rintveld Altrecht Mental Health Institute Zeist The Netherlands

Amplexa Genetics A S Odense Denmark

Amsterdam Public Health Research Institute VU Medical Center Amsterdam the Netherlands

Analytic Translational Genetics Unit Massachusetts General Hospital Harvard Medical School Boston Massachusetts USA

APC Microbiome Ireland University College Cork Cork Ireland

Arkin Amsterdam the Netherlands

Ashford and St Peters NHS Foundation Trust Surrey UK

Assistance Publique Hôpitaux de Paris Départment de Neurologie Groupe Hospitalier Pitié Salpêtrière Paris France

Assistance Publique Hopitaux de Paris GH Pitié Salpêtrière Paris France

Austin Health Heidelberg Australia

Avera Institute for Human Genetics Sioux Falls SD USA

AXA Research Fund and Sorbonne University Chair Paris France

Banner Sun Health Research Institute Sun City AZ USA

Bela Menso Brain and Behaviour Centre James Cook University Varsity Lakes Australia

Big Data Institute Li Ka Shing Centre for Health Information and Discovery University of Oxford Oxford UK

Bio4Dreams Srl Milan Italy

Bioinformatics Interdepartmental Program University of California Los Angeles Los Angeles CA USA

Bioinformatics Research Centre Aarhus University Aarhus Denmark

Biomedical Network Research Centre on Mental Health Instituto de Salud Carlos 3 Madrid Spain

Biopsychosocial Corporation Vienna Austria

BioRealm Walnut California USA

Biosystems and Integrative Sciences Institute BioISI University of Lisboa Lisboa Portugal

Black Dog Institute Sydney Australia

Bond University Faculty of Society and Design Robina Australia

Boston University School of Medicine Boston MA USA

Boston VA Research Institute Boston MA USA

Brain and Mind Centre University of Sydney Sydney Australia

Brain and Spine Institute INSERM U 1127 CNRS UMR 7225 Paris France

Brain Center Rudolf Magnus University Medical Center Utrecht Utrecht The Netherlands

Brigham and Women's Hospital Boston MA

Brigham Women's Hospital Division of Aging Harvard Medical School Boston MA USA

Broad Institute of MIT and Harvard Cambridge USA

Butler Hospital Providence RI USA

Byrd Alzheimer's Institute University of South Florida Tampa FL USA

Cambridge Institute for Medical Research University of Cambridge Cambridge UK

Cardiff University Medical Research Council Center for Neuropsychiatric Genetics and Genomics Institute of Psychology Medicine and Clinical Neuroscience Cardiff UK

Cardiovascular Health Research Unit Department of Medicine University of Washington Seattle WA USA

Carver College of Medicine University of Iowa Iowa City IA USA

Center for Applied Genomics of The Children's Hospital of Philadelphia Philadelphia PA USA

Center For Autism Research and Treatment Semel Institute David Geffen School of Medicine University of California Los Angeles Los Angeles CA USA

Center for Genomic Medicine Kyoto University Graduate School of Medicine Kyoto Japan

Center for Genomic Medicine Massachusetts General Hospital Boston MA USA

Center for Integrative Genomics University of Lausanne Lausanne Switzerland

Center for Integrative Sequencing Aarhus University Aarhus Denmark

Center for Molecular Medicine Karolinska University Hospital Stockholm Sweden

Center for Molecular Medicine University Medical Center Utrecht Utrecht the Netherlands

Center for Neurobehavioral Genetics Semel Institute for Neuroscience and Human Behavior University of California at Los Angeles Los Angeles CA USA

Center for Psychiatric Genetics NorthShore University HealthSystem Research Institute Evanston IL USA

Center for Psychiatric Research Maine Medical Center Research Institute Portland ME USA

Center for Translational and Computational Neuroimmunology Columbia University Medical Center New York NY USA

Center of Mental Health Department of Child and Adolescent Psychiatry Psychosomatics and Psychotherapy University Hospital of Wuerzburg Wuerzburg Germany

Central Institute of Mental Health Department of Genetic Epidemiology in Psychiatry Medical Faculty Mannheim University of Heidelberg Mannheim Germany

Centre d'Etudes et de Recherches en Psychopathologie et Psychologie de la Santé Université Toulouse Jean Jaurès Toulouse France

Centre de Recherche du Centre Hospitalier de l'Universite de Montreal and Department of Neurosciences Université de Montréal Montréal Canada

Centre for Genetic Epidemiology Institute for Clinical Epidemiology and Applied Biometery University of Tubingen Germany

Centre for Genomic and Experimental Medicine University of Edinburgh Edinburgh UK

Centre for Genomic Regulation Barcelona Spain

Centre for Genomic Sciences University of Hong Kong Hong Kong

Centre for Neural Engineering Department of Electrical and Electronic Engineering University of Melbourne Victoria Australia

Centre for Neuroimaging and Cognitive Genomics National University of Ireland Galway Galway Ireland

Centre for Prevention of Stroke and Dementia Nuffield Department of Clinical Neurosciences University of Oxford Oxford UK

Centre for Psychiatry Barts and the London School of Medicine and Dentistry London UK

Centre for Public Health Queens University Belfast Belfast UK

Centre Hosp Univ Lille Lille France

Centre National de Recherche en Génomique Humaine Institut de biologie François Jacob CEA Evry France

Centre National Maladie 'Syndrome Rare Gilles de la Tourette' Groupe Hospitalier Pitié Salpêtrière Paris France

Centro de Investigación Biomédica en Red de Enfermedades Raras Instituto de Salud Carlos 3 Madrid Spain

Centro de Investigación Biomédica en Red sobre Enfermedades Neurodegenerativas Madrid Spain

CERESA Toulouse France

Chalfont Centre for Epilepsy Bucks UK

Charite Universitatsmedizin Berlin Berlin Germany

Child and Adolescent Mental Health Centre Mental Health Services Capital Region Copenhagen University of Copenhagen Copenhagen Denmark

Child and Adolescent Mental Health Unit Hospital Universitario Mútua de Terrassa Barcelona Spain

Child and Adolescent Psychiatry and Psychotherapy Central Institute of Mental Health Medical Faculty Mannheim University of Heidelberg Mannheim Germany

Child and Adolescent Psychiatry Department Robert Debre Hospital APHP Paris France

Child and Adolescent Psychiatry Psychotherapy University Medical Center Goettingen Germany

Child and Adolescent Psychiatry Unit Department of Psychiatry Federal University of São Paulo Brazil

Child and Youth Mental Health Service Children's Health Queensland Health and Hospital Service Brisbane Australia

Child Developmental Center Hospital Pediátrico Centro Hospitalar e Universitário de Coimbra Coimbra Portugal

Child Health Research Centre University of Queensland Brisbane Australia

Children's Hospital Los Angeles Los Angeles CA USA

Children's Hospital of Philadelphia Philadelphia PA USA

Chinese University of Hong Kong Hong Kong

Clinic for Child and Adolescent Psychiatry and Psychotherapy University of Duisburg Essen Essen Germany

Clinic of Internal Medicine 1 University Hospital Schleswig Holstein Kiel Germany

Clinica Galatea y PAIMM Mental Health Program for Impaired Physicians Barcelona Spain

Clinical Research Grupo Médico Carracci Mexico City Mexico

Clinicum Department of Public Health University of Helsinki Finland

Cognitive Genetics and Cognitive Therapy Group Neuroimaging Cognition and Genomics Centre School of Psychology and Discipline of Biochemistry National University of Ireland Galway Galway Ireland

Cold Spring Harbor Laboratory Cold Spring Harbor NY USA

College of Medicine University of Kentucky Lexington KY USA

Cologne Center for Genomics University of Cologne Cologne Germany

Columbia University New York NY USA

Communication and Research Unit for Musculoskeletal Disorders Oslo University Hospital Oslo Norway

Computational Research Division Lawrence Berkeley National Laboratory University of California at Berkeley Berkeley CA USA

Cooper Medical School of Rowan University Camden NJ USA

CWZ Hospital Nijmegen Netherlands

Danish Epilepsy Centre Filadelfia Dianalund Denmark

Danish Headache Center Rigshospitalet Glostrup and University of Copenhagen Copenhagen Denmark

De Bascule Amsterdam The Netherlands

Dementia Research Centre UCL Institute of Neurology London UK

Dementia Research Institute Cardiff University Cardiff UK

Departamento de Genética Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

Department Clinical Genetics VU University Medical Center Amsterdam Neuroscience Amsterdam The Netherlands

Department Complex Trait Genetics Center for Neurogenomics and Cognitive Research VU University Amsterdam The Netherlands

Department for Congenital Disorders Statens Serum Institut Copenhagen Denmark

Department of Adolescent Psychiatry Helsinki University Central Hospital Helsinki Finland

Department of Adult Psychiatry Poznan University of Medical Sciences Poland

Department of Behavioral Sciences Rush Medical College Chicago IL USA

Department of Biochemistry and Molecular Biology 2 Institute of Neurosciences Center for Biomedical Research University of Granada Granada Spain

Department of Biochemistry and Molecular Biology Indiana University School of Medicine Indianapolis IN USA

Department of Biological and Medical Psychology University of Bergen Bergen Norway

Department of Biological Psychology Vrije Universiteit Amsterdam Amsterdam The Netherlands

Department of Biological Sciences Purdue University West Lafayette Indiana USA

Department of Biology Brigham Young University Provo UT USA

Department of Biomedical Sciences Cooper Medical School of Rowan University Camden NJ USA

Department of Biomedicine Aarhus University Aarhus Denmark

Department of Biomedicine and Human Genetics Aarhus University Aarhus Denmark

Department of Biomedicine University of Basel Basel Switzerland

Department of Biostatistics University of Washington Seattle WA USA

Department of Cancer Epidemiology and Prevention Cancer Center and M Sklodowska Curie Institute of Oncology Warsaw Poland

Department of Cell Biology SUNY Downstate Medical Center Brooklyn NY USA

Department of Cerebrovascular Diseases Fondazione IRCCS Istituto Neurologico Carlo Besta Milano Italy

Department of Child and Adolescent Psychiatry Academic Medical Center University of Amsterdam Amsterdam the Netherlands

Department of Child and Adolescent Psychiatry and Psychosomatic Medicine of University Clinics RWTH Aachen Aachen Germany

Department of Child and Adolescent Psychiatry Faculty of Medicine Technischen Universität Dresden Dresden Germany

Department of Child and Adolescent Psychiatry Psychosomatics and Psychotherapy University Hospital Frankfurt Goethe University Frankfurt am Main Germany

Department of Child and Adolescent Psychiatry University Medical Center Groningen University of Groningen Groningen The Netherlands

Department of Child Psychiatry Medical University of Warsaw Warsaw Poland

Department of Clinical and Experimental Epilepsy NIHR University College London Hospitals Biomedical Research Centre UCL Institute of Neurology London

Department of Clinical and Experimental Epilepsy UCL Institute of Neurology London UK

Department of Clinical Chemistry Fimlab Laboratories and Finnish Cardiovascular Research Center Tampere Faculty of Medicine and Life Sciences University of Tampere Tampere Finland

Department of Clinical Neuroscience Centre for Psychiatry Research Karolinska Institutet Stockholm Sweden

Department of Clinical Neurosciences University of Cambridge Cambridge UK

Department of Clinical Sciences University of Copenhagen Copenhagen Denmark

Department of Cognitive Neuroscience Donders Institute for Brain Cognition and Behavior Radboud University Medical Centre Nijmegen The Netherlands

Department of Computer Science New Jersey Institute of Technology New Jersey USA

Department of Environmental Health Harvard T H Chan School of Public Health Boston MA USA

Department of Epidemiology Erasmus Medical Centre Rotterdam the Netherlands

Department of Epidemiology Harvard T H Chan School of Public Health Boston Massachusetts USA

Department of Epileptology University Bonn Medical Center Bonn Germany

Department of Epileptology University Hospital Bonn Bonn Germany

Department of Genetics and Genomic Sciences and Institute for Genomics and Multiscale Biology Icahn School of Medicine at Mount Sinai New York NY USA

Department of Genetics and Psychiatry University of North Carolina School of Medicine Chapel Hill North Carolina USA

Department of Genetics Center for Molecular Medicine University Medical Center Utrecht Utrecht The Netherlands

Department of Genetics Microbiology and Statistics Faculty of Biology University of Barcelona Barcelona Spain

Department of Genetics University Medical Center Utrecht the Netherlands

Department of Genetics Yale University School of Medicine New Haven CT USA

Department of Genomics Life and Brain Center University of Bonn Bonn Germany

Department of Gerontology Hôpital Broca AH HP Paris France

Department of Human Genetics CHU Sart Tilman University of Liège Liège Belgium

Department of Human Genetics David Geffen School of Medicine University of California at Los Angeles Los Angeles CA USA

Department of Human Genetics Institute of Molecular Genetics Russian Academy of Sciences Moscow Russia

Department of Human Genetics Leiden University Medical Center Leiden The Netherlands

Department of Human Genetics Radboud University Medical Center Nijmegen The Netherlands

Department of Human Genetics Wellcome Sanger Institute Hinxton Cambridgeshire UK

Department of Mathematics Massachusetts Institute of Technology Cambridge Massachusetts USA

Department of Medical Epidemiology and Biostatistics Karolinska Institutet Stockholm Sweden

Department of Medical Genetics Cambridge Institute for Medical Research Cambridge UK

Department of Medical Genetics Haukeland University Hospital Bergen Norway

Department of Medical Genetics Medical University Sofia Bulgaria

Department of Medical Informatics and Clinical Epidemiology Oregon Health and Science University Portland OR USA

Department of Medicine Boston University School of Medicine Boston MA USA

Department of Medicine Harvard Medical School Boston MA USA

Department of Medicine Surgery and Dentistry Scuola Medica Salernitana University of Salerno Italy

Department of Medicine The University of Melbourne Austin Health Melbourne Victoria Australia

Department of Medicine University of Washington WA USA

Department of Mental and Physical Health and Preventive Medicine University of Campania luigi Vanvitelli Naples Italy

Department of Mental Health Johns Hopkins Bloomberg School of Public Health Baltimore MD USA

Department of Molecular and Cellular Therapeutics Royal College of Surgeons in Ireland Dublin Ireland

Department of Molecular Bases of Human Genetics Institute of Molecular Genetics Russian Academy of Sciences Moscow Russia

Department of Molecular Biology and Genetics Democritus University of Thrace Alexandroupolis Greece

Department of Molecular Medicine and Surgery Karolinska Institutet Stockholm Sweden

Department of Molecular Neuroscience Institute of Neurology University College London London UK

Department of Neurodegeneration UCL Institute of Neurology London UK

Department of Neurological Sciences Rush Medical College Chicago IL USA

Department of Neurology 2 Kepler University Clinic Johannes Kepler University Linz Austria

Department of Neurology Admiraal De Ruyter Hospital Goes The Netherlands

Department of Neurology and Epileptology Hertie Institute for Clinical Brain Research University of Tuebingen Tuebingen Germany

Department of Neurology and Epileptology University Hospital Tuebingen Tuebingen Germany

Department of Neurology and Neurosurgery McGill University Montréal Canada

Department of Neurology and Parkinson's Disease Research Education and Clinical Care Center Portland Veterans Affairs Medical Center Portland OR USA

Department of Neurology Antwerp University Hospital Antwerp Belgium

Department of Neurology Beth Israel Deaconess Medical Center Boston MA USA

Department of Neurology Brigham and Women's Hospital Harvard Medical School Boston MA USA

Department of Neurology College of Medicine Jacksonville University of Florida Jacksonville FL USA

Department of Neurology Duke University School of Medicine Durham NC USA

Department of Neurology Helsinki University Central Hospital Helsinki Finland

Department of Neurology Hôpital Erasme Université Libre de Bruxelles Brussels Belgium

Department of Neurology Inselspital Bern University Hospital University of Bern Switzerland

Department of Neurology Jagiellonian University Medical College Kraków Poland

Department of Neurology Klinikum rechts der Isar Technical University of Munich Munich Germany

Department of Neurology Leiden University Medical Center Leiden The Netherlands

Department of Neurology Massachusetts General Hospital Boston MA USA

Department of Neurology Medical University of Vienna Austria

Department of Neurology Oregon Health and Science University Portland OR USA

Department of Neurology Oslo University Hospital Oslo Norway

Department of Neurology School of Medicine Emory University Atlanta GA USA

Department of Neurology Thomas Jefferson University Philadelphia PA USA

Department of Neurology University Hospital Essen Germany

Department of Neurology University of California San Francisco CA USA

Department of Neurology University of Cincinnati Cincinnati OH USA

Department of Neurology University of Florida Gainesville FL USA

Department of Neurology University of Munich Hospital Grosshadern University of Munich Germany

Department of Neurology University of Ulm Ulm Germany

Department of Neurology Yale School of Medicine New Haven CT USA

Department of Neuroscience Icahn School of Medicine at Mount Sinai New York NY USA

Department of Neuroscience Karolinska Institutet Stockholm Sweden

Department of Ophthalmology University of Helsinki and Helsinki University Hospital Helsinki Finland

Department of Paediatrics Royal Children's Hospital The University of Melbourne Melbourne Victoria Australia

Department of Pathology and Laboratory Medicine Perelman School of Medicine University of Pennsylvania Philadelphia PA USA

Department of Pathology Duke University School of Medicine Durham NC USA

Department of Pathology University of Michigan Medical School Ann Arbor MI USA

Department of Pathology University of Utah Salt Lake City UT USA

Department of Pathology University of Washington Seattle WA USA

Department of Pediatrics Nemours A 1 duPont Hospital for Children Wilmington DE USA

Department of Pediatrics The Perelman School of Medicine University of Pennsylvania Philadelphia PA USA

Department of Pharmacy and Biotechnology University of Bologna Bologna Italy

Department of Physiology Anatomy and Genetics University of Oxford Oxford UK

Department of Physiology Saint Antoine Hospital Paris France

Department of Psychiatry Academic Medical Center University of Amsterdam Amsterdam the Netherlands

Department of Psychiatry and Behavioral Neuroscience University of Chicago Chicago IL USA

Department of Psychiatry and Behavioral Science Stanford University Stanford California USA

Department of Psychiatry and Behavioral Sciences Duke University Durham NC USA

Department of Psychiatry and Behavioral Sciences Johns Hopkins University Baltimore MD USA

Department of Psychiatry and Behavioral Sciences NorthShore University HealthSystem Research Institute Evanston IL USA

Department of Psychiatry and Biobehavioral Sciences Semel Institute for Neuroscience and Human Behavior David Geffen School of Medicine University of California at Los Angeles Los Angeles CA USA

Department of Psychiatry and Global Brain Health Institute Trinity College Dublin Ireland

Department of Psychiatry and Human Behavior University of California Irvine Irvine CA USA

Department of Psychiatry and Human Genetics Donders Institute for Brain Cognition and Behaviour Radboud University Medical Center Nijmegen The Netherlands

Department of Psychiatry and Legal Medicine Universitat Autònoma de Barcelona Barcelona Spain

Department of Psychiatry and Psychotherapy Friedrich Alexander Universität Erlangen Nürnberg University Hospital Erlangen Germany

Department of Psychiatry and Psychotherapy Jena University Hospital Jena Germany

Department of Psychiatry and Psychotherapy Medical University of Vienna Vienna Austria

Department of Psychiatry and Psychotherapy University Hospital Carl Gustav Carus Technische Universität Dresden Dresden Germany

Department of Psychiatry and Psychotherapy University Medical Center Göttingen Göttingen Germany

Department of Psychiatry and Psychotherapy University Medicine Greifswald Greifswald Germany

Department of Psychiatry and UF Genetics Institute University of Florida Gainesville FL USA

Department of Psychiatry Carver College of Medicine University of Iowa Iowa City IA USA

Department of Psychiatry Center for Neurobiology and Behavior Perelman School of Medicine at the University of Pennsylvania Philadelphia PA USA

Department of Psychiatry Dalhousie University Halifax Canada

Department of Psychiatry David Geffen School of Medicine University of California at Los Angeles Los Angeles CA USA

Department of Psychiatry Faculdade de Medicina Universidade Federal do Rio Grande do Sul Porto Alegre Brazil

Department of Psychiatry GGZ INGEEST Amsterdam the Netherlands

Department of Psychiatry Harvard Medical School Boston MA USA

Department of Psychiatry Hospital for Sick Children and University of Toronto Toronto Canada

Department of Psychiatry Hospital Universitari Vall d'Hebron Barcelona Spain

Department of Psychiatry Icahn School of Medicine at Mount Sinai New York NY USA

Department of Psychiatry McGill University Montréal Canada

Department of Psychiatry Psychosomatic Medicine and Psychotherapy University Hospital Frankfurt Frankfurt am Main Germany

Department of Psychiatry Psychosomatics and Psychotherapy Center of Mental Health University Hospital Wuerzburg Wuerzburg Germany

Department of Psychiatry Radboud University Medical Center Nijmegen The Netherlands

Department of Psychiatry Royal College of Surgeons in Ireland Dublin Ireland

Department of Psychiatry Saarland University Hospital Homburg Germany

Department of Psychiatry Sao Paulo Medical School University of Sao Paulo Sao Paulo Brazil

Department of Psychiatry Sidney Kimmel Medical College Thomas Jefferson University Philadelphia PA USA

Department of Psychiatry Tufts University School of Medicine Boston MA USA

Department of Psychiatry University College Cork Cork Ireland

Department of Psychiatry University of Basel Basel Switzerland

Department of Psychiatry University of British Columbia Vancouver Canada

Department of Psychiatry University of California Los Angeles Los Angeles CA USA

Department of Psychiatry University of California San Diego La Jolla CA USA

Department of Psychiatry University of California San Francisco San Francisco CA USA

Department of Psychiatry University of Groningen and University Medical Center Groningen the Netherlands

Department of Psychiatry University of Helsinki Helsinki Finland

Department of Psychiatry University of Illinois at Chicago Chicago IL USA

Department of Psychiatry University of Medical Sciences Poznan Poland

Department of Psychiatry University of Michigan Ann Arbor MI USA

Department of Psychiatry University of North Carolina at Chapel Hill Chapel Hill NC USA

Department of Psychiatry University of Oxford Oxford UK

Department of Psychiatry University of Perugia Perugia Italy

Department of Psychiatry University of Toronto Toronto Canada

Department of Psychiatry Virginia Commonwealth University Richmond VA USA

Department of Psychiatry VU University Medical Center Amsterdam The Netherlands

Department of Psychiatry Yale School of Medicine New Haven CT USA

Department of Psychological and Brain Sciences University of Iowa Iowa City IA USA

Department of Psychology Emory University Atlanta GA USA

Department of Psychology University of Denver Denver CO USA

Department of Psychology University of Oslo Norway

Department of Psychosis Studies Institute of Psychiatry King's College London London UK

Department of Public Health University of Helsinki Helsinki Finland

Department of Radiology and Nuclear Medicine Erasmus Medical Centre Rotterdam the Netherlands

Department of Research and Innovation GGZ Ingeest Specialized Mental Health Care Amsterdam the Netherlands

Department of Rheumatology CHU Sart Tilman University of Liège Liège Belgium

Department of Specialized Trainings GGZ Drenthe Mental Health Care Services Assen the Netherlands

Department of Statistics and Applied Probability University of California Santa Barbara CA USA

Department of Translational Psychiatry School for Mental Health and Neuroscience Maastricht University Maastricht The Netherlands

Department of Translational Research in Psychiatry Max Planck Institute of Psychiatry Munich Germany

Departments of Medicine and Psychiatry School of Medicine University of Cantabria IDIVAL University Hospital Marqués de Valdecilla Santander Spain

Departments of Neurology and Neurosurgery University of Florida Gainesville FL USA

Departments of Pediatrics and Neurology Ohio State University Columbus OH USA

Departments of Psychiatry and Genetics Washington University School of Medicine St Louis MO USA

Departments of Psychiatry and Medical Genetics Cumming School of Medicine University of Calgary Calgary Canada

Departments of Psychiatry and Nutrition University of North Carolina at Chapel Hill Chapel Hill NC USA

Departments of Psychiatry and of Neuroscience and Physiology SUNY Upstate Medical University Syracuse NY USA

Departments of Psychology and Human and Molecular Genetics College Behavioral and Emotional Health Institute Virginia Commonwealth University Richmond Virginia

Dept of Clinical Genetics Our Lady's Children's Hospital Crumlin Dublin Ireland

Dept of Medicine University of Maryland School of Medicine Baltimore MD USA

Depto Farmacogenética Instituto Nacional de Psiquiatria Ramon de la Fuente Muñiz Mexico City Mexico

Discipline of Psychiatry University of Adelaide Adelaide Australia

Division Biomedical Genetics University Medical Center Utrecht Utrecht the Netherlands

Division of Adolescent and Child Psychiatry Department of Psychiatry Lausanne University Hospital Lausanne Switzerland

Division of Child and Adolescent Psychiatry Department of Psychiatry and Human Behavior Brown University Providence RI USA

Division of Clinical Neuroscience Oslo University Hospital and University of Oslo Oslo Norway

Division of Cognitive and Behavioral Neurology Brigham and Women's Hospital Boston MA USA

Division of Human Genetics Children's Hospital of Philadelphia Philadelphia PA USA

Division of Life Science Hong Kong University of Science and Technology Hong Kong Special Administrative Region China

Division of Medical Genetics Department of Medicine University of Washington Seattle WA USA

Division of Mental Health and Addiction Oslo University Hospital Oslo Norway

Division of Molecular Genome Analysis and Division of Cancer Genome Research German Cancer Research Center Heidelberg Germany

Division of Molecular Psychiatry Center of Mental Health University of Wuerzburg Wuerzburg Germany

Division of Neurocritical Care and Emergency Neurology Massachusetts General Hospital Boston MA USA

Division of Neurology Children's Hospital of Philadelphia Philadelphia PA USA

Division of Neuropsychiatry University College London London UK

Division of Neuroscience School of Medicine University of Dundee Dundee UK

Division of Preventive Medicine Brigham and Women's Hospital Boston MA USA

Division of Psychiatry Centre for Clinical Brain Sciences University of Edinburgh Edinburgh UK

Division of Psychiatry Haukeland University Hospital Bergen Norway

Division of Psychiatry Molecular Psychiatry Laboratory University College London London UK

Division of Psychiatry University of Edinburgh Edinburgh UK

Division of Psychological and Social Medicine and Developmental Neurosciences Faculty of Medicine Technischen Universität Dresden Dresden Germany

Donders Institute for Brain Cognition and Behaviour Radboud University Nijmegen The Netherlands

Douglas Mental Health University Institute McGill University Montreal Canada

Eating Disorders Unit 1st Psychiatric Department National and Kapodistrian University of Athens Athens Greece

Eating Disorders Unit Department of Child and Adolescent Psychiatry Medical University of Vienna Vienna Austria

Eating Disorders Unit Department of Psychiatry 1st Faculty of Medicine Charles University Prague Czech Republic

Epilepsy Center Neurocenter Kuopio University Hospital Kuopio Finland

Epilepsy Foundation in the Netherlands Heemstede the Netherlands

Epilepsy Research Centre University of Melbourne Austin Health Heidelberg Australia

Epilepsy Research Centre University of Melbourne Heidelberg Australia

Epilepsy Society Chalfont St Peter Bucks UK

Escuela de Ciencias de la Salud Universidad Pontificia Bolivariana Medellín Colombia

Estonian Genome Center Institute of Genomics University of Tartu Tartu Estonia

Faculdade de Medicina FMUSP Universidade de São Paulo São Paulo Brazil

Facultat de Medicina i Ciències de la Salut Universitat Internacional de Catalunya Barcelona Spain

Faculté de Médecine Université Paris Est Créteil France

Faculty of Health and Medicine University of Newcastle Callaghan Australia

Faculty of Science Medicine and Health University of Wollongong Wollongong Australia

Faculty of Social Sciences University of Utrecht Utrecht the Netherlands

Fixel Center for Neurological Diseases University of Florida Gainesville FL USA

Florey Institute of Neuroscience and Mental Health Melbourne Victoria Australia

Fondation FondaMental Créteil France

Framingham Heart Study Framingham MA USA

Frederick W Thompson Anxiety Disorders Centre Sunnybrook Health Sciences Centre Toronto Canada

Functional Genomics Center Zurich ETH UZH Zurich Zurich Switzerland

Fundació ACE Institut Català de Neurociències Aplicades Barcelona Spain and Universitat Internacional de Catalunya Barcelona Spain

Fundació Docència i Recerca Mútua Terrassa

General University Hospital Prague Czech Republic

Genes and Disease Programme Centre for Genomic Regulation Barcelona Spain

Genomics Medicine Ireland Dublin Ireland

Geriatrics Research and Education Clinical Center Baltimore Veterans Administration Medical Center Baltimore MD USA

German Center for Neurodegenerative Diseases Tübingen Germany

German Centre for Cardiovascular Research Partner Site Greifswald Greifswald Germany

Gertrude H Sergievsky Center and Dept of Neurology Columbia University New York NY USA

Glenn Biggs Institute for Alzheimer's and Neurodegenerative Diseases University of Texas Health Sciences Center San Antonio Texas USA

Greek Association of Alzheimer's Disease and Related Disorders Thessaloniki Greece

Groote Schuur Hospital Cape Town South Africa

Grupo de Genética Molecular Instituto de Biología Facultad de Ciencias Exactas y Naturales Universidad de Antioquia Medellín Colombia

Harvard Medical School Boston MA USA

Harvard Medical School McLean Hospital Belmont MA USA

Headache Research Group VHIR Autonomous University of Barcelona Barcelona Spain

Headache Unit Neurology Department Hospital Vall d'Hebron Barcelona Spain

Hertie Institute for Clinical Brain Research University of Tübingen Tübingen Germany

Hong Kong Sanatorium and Hospital Hong Kong

Hôpital Paul Brousse Université Paris Sud XI Le Kremlin Bicêtre Paris France

Hospital for Psychiatry and Psychotherapy Cologne Germany

Hospital for Sick Children Toronto Canada

Hospital of Telemark Kragerø Norway

Human Genetics and Cognitive Functions Institut Pasteur Paris France

Human Genetics Branch National Institute of Mental Health National Institutes of Health US Department of Health and Human Services Bethesda MD USA

Human Genetics School of Life Sciences University of Nottingham Nottingham UK

Human Genomics Research Group Department of Biomedicine University of Basel Basel Switzerland

Hunter Medical Research Institute Newcastle Australia

Hunter New England Local Health District Newcastle Australia

Icahn School of Medicine at Mount Sinai New York NY USA

IGBMC CNRS UMR 7104 INSERM U964 Université de Strasbourg Illkirch France

Illumina Inc San Diego CA USA

Indiana Alzheimer Disease Center Indiana University School of Medicine Indianapolis IN USA

INSERM U 1127 CNRS UMR 7225 Sorbonne Universités UPMC Univ Paris 06 UMR S 1127 Paris France

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INSERM U1220 IRSD Toulouse France

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Inserm U955 Psychiatrie Translationnelle Créteil France

INSERM UMR 1219 Bordeaux France

Institut de Biomedicina de la Universitat de Barcelona Barcelona Spain

Institut de Recerca Sant Joan de Déu Barcelona Spain

Institut Pasteur de Lille U1167 Lille France

Institut Universitaire de France Paris France

Institute for Genomic Medicine Columbia University Medical Center New York NY USA

Institute for Human Genetics University of California San Francisco San Francisco CA USA

Institute for Memory Impairments and Neurological Disorders University of California Irvine CA USA

Institute for Molecular Bioscience University of Queensland Brisbane Australia

Institute for Molecular Medicine Finland HiLIFE University of Helsinki Helsinki Finland

Institute for Precision Health University of California Los Angeles Los Angeles CA USA

Institute for Regional Health Services University of Southern Denmark Odense Denmark

Institute for Stroke and Dementia Research Klinikum der Universitaet Muenchen Munich Germany

Institute for Stroke and Dementia Research University Hospital LMU Munich Munich Germany

Institute for Translational Genomics and Population Sciences Departments of Pediatrics and Medicine LABioMed at Harbor UCLA Medical Center Torrance CA USA

Institute of Applied Health Research University of Birmingham UK

Institute of Biological Psychiatry Roskilde Denmark

Institute of Cardiovascular Research Royal Holloway University of London London UK

Institute of Clinical Chemistry and Laboratory Medicine University Medicine Greifswald Greifswald Germany

Institute of Clinical Medicine School of Medicine Faculty of Health Sciences University of Eastern Finland Kuopio Finland

Institute of Clinical Molecular Biology Kiel University and University Hospital Schleswig Holstein Kiel Germany

Institute of Environmental Medicine Karolinska Institutet Stockholm Sweden

Institute of Experimental Epileptology and Cognition Research University Bonn Medical Center Bonn Germany

Institute of Human Genetics School of Medicine University of Bonn and University Hospital Bonn Bonn Germany

Institute of Human Genetics University Hospital Essen University Duisburg Essen Essen Germany

Institute of Human Genetics University Medical Center Hamburg Eppendorf Hamburg Germany

Institute of Human Genetics University of Bonn Bonn Germany

Institute of Human Genetics University of Ulm Ulm Germany

Institute of Medical Genetics and Pathology University Hospital Basel Basel Switzerland

Institute of Medical Informatics and Statistics Kiel University Kiel Germany

Institute of Memory and Alzheimer's Disease Department of Neurology Pitié Salpêtrière Hospital AP HP Paris France

Institute of Mental Health Lee Kong Chian School of Medicine Nanyang Technological University Singapore Singapore

Institute of Mental Health Singapore Singapore

Institute of Mental Health University of British Columbia Vancouver Canada

Institute of Mental Health University of Nottingham Nottingham UK

Institute of Neuropathology Bonn University Medical School Bonn Germany

Institute of Neuroscience and Medicine Research Centre Jülich Jülich Germany

Institute of Neuroscience Newcastle University Newcastle UK

Institute of Prion Diseases and MRC Prion Unit University College London London UK

Institute of Psychiatric Phenomics and Genomics University Hospital LMU Munich Munich Germany

Institute of Psychiatry Psychology and Neuroscience King's College London London UK

Institute of Psychiatry Psychology and Neuroscience Kings College London UK

Institute of Psychiatry Psychology and Neuroscience Social Genetics and Developmental Psychiatry Center MRC Kings College London London UK

Institute of Psychological Medicine and Clinical Neurosciences School of Medicine Cardiff University Cardiff UK

Institute of Public Health Charité Universitätsmedizin Berlin Berlin Germany

Institute of Social Medicine Occupational Health and Public Health University of Leipzig Leipzig Germany

Institute of Translational Medicine University of Liverpool Liverpool UK

Instituto de Investigación Biomédica de Málaga Malaga Spain

Instituto Nacional de Saúde Doutor Ricardo Jorge Lisboa Portugal

Interfaculty Institute for Genetics and Functional Genomics University Medicine Greifswald Greifswald Germany

Janssen Research and Development LLC Titusville NJ USA

Johns Hopkins University School of Medicine Baltimore MD USA

K G Jebsen Centre for Neuropsychiatric Disorders Department of Biomedicine University of Bergen Norway

K G Jebsen Centre for Neuropsychiatric Disorders Department of Clinical Science University of Bergen Norway

K G Jebsen Centre for Research on Neuropsychiatric Disorders University of Bergen Bergen Norway

Kaiser Permanente Washington Health Research Institute Seattle WA USA

Karakter Child and Adolescent Psychiatry University Center Nijmegen The Netherlands

Kartini Clinic Portland OR USA

Keck School of Medicine of the University of Southern California Los Angeles CA USA

Kiel Pain and Headache Center Kiel Germany

King's College Hospital London UK

KIZ CUHK Joint Laboratory of Bioresources and Molecular Research of Common Diseases Kunming Institute of Zoology Chinese Academy of Sciences Kunming China

Krembil Research Institute University Health Network Toronto Canada

Laboratorio de Genética Molecular SIU Universidad de Antioquia Medellín Colombia

Laboratory for Statistical Analysis RIKEN Center for Integrative Medical Sciences Yokohama Japan

Laboratory of Neurogenetics G Gaslini Institute Genova Italy

Laboratory of Neurogenetics National Institute on Aging National Institutes of Health Bethesda MD USA

Laboratory of Pharmaceutical Biotechnology Ghent University Ghent Belgium

Laboratory of Psychiatric Genetics Department of Psychiatry Poznan University of Medical Sciences Poznan Poland

Laboratory of Psychiatric Neurobiology Institute of Molecular Medicine 1 M Sechenov 1st Moscow State Medical University Moscow Russia

Landspitali National University Hospital Reykjavik Iceland

Latvian Biomedical Research and Study Centre Riga Latvia

Layton Aging and Alzheimer's Disease Center Oregon Health and Science University Portland OR USA

Lee Kong Chian School of Medicine Nanyang Technological University Singapore Singapore

LifeOmic Indianapolis IN USA

Lou Ruvo Center for Brain Health Neurological Institute Cleveland Clinic Cleveland OH USA

Lundbeck Foundation Initiative for Integrative Psychiatric Research iPSYCH Aarhus Denmark

Maastricht University Medical Centre Maastricht the Netherlands

Massachusetts General Hospital Boston MA USA

Mathison Centre for Mental Health Research and Education Hotchkiss Brain Institute Cumming School of Medicine University of Calgary Calgary Canada

Maurice Wohl Clinical Neuroscience Institute Department of Basic and Clinical Neuroscience King's College London London UK

Maurice Wohl Clinical Neuroscience Institute Institute of Psychiatry Psychology and Neuroscience King's College London London UK

Max Planck Institute of Psychiatry Munich Germany

McKnight Brain Institute University of Florida Gainesville FL USA

McKusick Nathans Institute of Genetic Medicine Johns Hopkins University School of Medicine Baltimore MD USA

McLaughlin Centre and Department of Molecular Genetics University of Toronto Toronto Canada

Me Inc Mountain View CA US

Medical Genetics Unit Department SDB University of Padova Padova Italy

Medical University of Vienna Austria

Melbourne Neuropsychiatry Centre Department of Psychiatry University of Melbourne and Melbourne Health Victoria Australia

Memory Research and Resources Center Department of Neurology Montpellier University Hospital Gui de Chauliac Montpellier France

Mindich Child Health and Development Institute Icahn School of Medicine at Mount Sinai New York NY USA

Minerva Neurosciences Inc Waltham MA USA

Molecular and Behavioral Neuroscience Institute University of Michigan Ann Arbor MI USA

Montreal Neurological Institute McGill University Montréal Canada

Moscow Institute of Physics and Technology Dolgoprudny Institusky 9 Moscow Russia

MRC Centre for Neuropsychiatric Genetics and Genomics Cardiff University Cardiff UK

MRC Integrative Epidemiology Unit and Bristol Medical School University of Bristol Bristol UK

MRC Social Genetic and Developmental Psychiatry Centre King's College London London UK

MRC Unit on Risk and Resilience in Mental Disorders Department of Psychiatry University of Cape Town Cape Town South Africa

Munich Cluster for Systems Neurology Munich Germany

National Alzheimer Coordinating Center Department of Epidemiology University of Washington Seattle WA USA

National Centre for Mental Health MRC Centre for Neuropsychiatric Genetics and Genomics Cardiff University Cardiff UK

National Centre for Register based Research Aarhus University Aarhus Denmark

National Institute for Health and Welfare Helsinki Finland

National Institute of Genomic Medicine Ciudad de México Mexico

National Institute of Mental Health Klecany Czech Republic

Nationwide Children's Hospital Columbus OH USA

NCBES Galway Neuroscience Centre National University of Ireland Galway Galway Ireland

Netherlands Institute for Neuroscience Royal Netherlands Academy of Arts and Sciences Amsterdam the Netherlands

Netherlands Twin Register Vrije Universiteit Amsterdam the Netherlands

Neurogenetics Group VIB CMN Antwerp Belgium

Neurogenetics Program Departments of Neurology and Human Genetics David Geffen School of Medicine University of California Los Angeles Los Angeles CA USA

Neuroimaging Cognition and Genomics Centre School of Psychology National University of Ireland Galway Galway Ireland

Neurology and Neurogenetics Core Framingham Heart Study Framingham MA USA

Neuropsychiatric Genetics Research Group Department of Psychiatry Trinity College Dublin Dublin Ireland

Neuropsychiatric Research Institute Fargo ND USA

Neuroscience Research Australia Sydney Australia

Neurosciences and Mental Health Program Research Institute Hospital for Sick Children Toronto Canada

Newcastle Upon Tyne Hospitals NHS Foundation Trust Newcastle UK

NIHR Biomedical Research Centre in Mental Health Maudsley Hospital London UK

NIHR Maudsley Biomedical Research Centre South London and Maudsley NHS Trust and King's College London London UK

NORMENT Div of Mental Health and Addiction University of Oslo Oslo Norway

NORMENT K G Jebsen Center for Psychosis Research Department of Clinical Science University of Bergen Norway

NORMENT K G Jebsen Centre for Psychosis Research Institute of Clinical Medicine University of Oslo Oslo Norway

Northumberland Tyne and Wear NHS Foundation Trust Northumberland UK

Norwegian Institute of Public Health Oslo Norway

NSW Health Pathology Newcastle Australia

NYU School of Medicine New York NY USA

Oregon Research Institute Eugene OR USA

Oslo University Hospital Oslo Norway

Ospedale San Raffaele Milano Italy

Oxford Centre for Diabetes Endocrinology and Metabolism Nuffield Department of Medicine University of Oxford Oxford UK

Oxford Health NHS Foundation Trust Oxford UK

Pediatric Neurology and Muscular Diseases Unit Department of Neurosciences Rehabilitation Ophthalmology Genetics Maternal and Child Health University of Genoa G Gaslini Institute Genova Italy

Pediatric Neurology Research Group Vall d'Hebron Research Institute Autonomous University of Barcelona Barcelona Spain

Perelman School of Medicine at the University of Pennsylvania Philadelphia PA USA

Pfizer Inc New York NY USA

PhD Program in Neuroscience University Milano Bicocca Monza Italy

Philipps Universität Marburg and Marburg University Hospital UKGM Marburg Germany

Priority Research Centre for Brain and Mental Health Research Hunter Medical Research Institute Newcastle Australia

Priority Research Centre for Health Behaviour University of Newcastle Callaghan Australia

Program in Genetics and Genome Biology Hospital for Sick Children Toronto Canada

Program in Medical and Population Genetics Broad Institute of MIT and Harvard Cambridge Massachusetts USA

Programa Corporatiu Neurodevelopment Disorders along Life Span Institut Català de la Salut Barcelona Spain

Psychiatric Genetics Unit Group of Psychiatry Mental Health and Addiction Vall d'Hebron Research Institute Universitat Autònoma de Barcelona Barcelona Spain

QIMR Berghofer Medical Research Institute Brisbane Australia

Quantinuum Research LLC San Diego CA USA

Queensland Brain Institute University of Queensland Brisbane Australia

Reference center for Gilles de la Tourette syndrome Pitie Salpetriere Hospital Paris France

Research School of Behavioural and Cognitive Neurosciences Department of Psychiatry University Medical Center Groningen University of Groningen Groningen the Netherlands

Research Unit Sørlandet Hospital Kristiansand Norway

Rheumatology Unit Department of Medicine Karolinska Institutet and Karolinska University Hospital Solna Sweden

Rouen University Hospital Rouen France

Rush Alzheimers Disease Center Chicago IL USA

Sackler School of Medicine Tel Aviv University Tel Aviv Israel

Sanders Brown Center on Aging University of Kentucky Lexington KY USA

Schizophrenia Research Institute Sydney Australia

School of Biomedical Sciences and Pharmacy The University of Newcastle Callaghan Australia

School of Biomedical Sciences Chinese University of Hong Kong Shatin Hong Kong

School of Biotechnology Dublin City University Glasnevin Dublin Ireland

School of Medical Sciences University of New South Wales Sydney Australia

School of Medicine and Medical Science University College Dublin Dublin Ireland

School of Medicine and Public Health University of Newcastle Callaghan Australia

School of Medicine Boston University Boston MA USA

School of Medicine Cardiff University Cardiff UK

School of Medicine University of North Dakota Grand Forks ND USA

School of Nursing Louisiana State University Health Sciences Center New Orleans LA USA

School of Psychiatry University of New South Wales Sydney Australia

School of Psychology and Counselling Faculty of Health Institute of Health and Biomedical Innovation Queensland University of Technology Queensland Australia

School of Psychology Cardiff University UK

School of Psychology Flinders University Adelaide Australia

School of Public Health Boston University Boston MA USA

Seaver Autism Center for Research and Treatment Icahn School of Medicine at Mount Sinai New York NY USA

Sección de Neuropediatría Instituto de Biomedicina de Sevilla Hospital Universitario Virgen del Rocío CSIC Universidad de Sevilla Seville Spain

Section of Epileptology Department of Neurology University Hospital RWTH Aachen Aachen Germany

Semel Institute for Neuroscience and Human Behavior David Geffen School of Medicine University of California at Los Angeles Los Angeles CA USA

Sheba Medical Center Ramat Gan Israel

Sorbonne Université CNRS UMR 7222 Institut des Systèmes Intelligents et Robotiques Paris France

Sorbonne Université Faculty of Médecine Paris France

Sorbonne Université INSERM CNRS Neuroscience Paris Seine Institut de Biologie Paris Seine Paris France

Sorbonne Universités UPMC Université Paris 06 UMR S 1127 CNRS UMR 7225 ICM Paris France

Sorbonne University GRC n° 21 Alzheimer Precision Medicine AP HP Pitié Salpêtrière Hospital Paris France

SRH University Heidelberg Academy for Psychotherapy Heidelberg Germany

Stanley Center for Psychiatric Research Broad Institute of MIT and Harvard Cambridge Massachusetts USA

Statistical and Genomic Epidemiology Laboratory Institute of Health and Biomedical Innovation Queensland University of Technology Brisbane Queensland Australia

Stella Maris Clinical Research Institute for Child and Adolescent Neuropsychiatry Pisa Italy

SUNY Downstate Medical Center Brooklyn NY USA

Tanz Centre for Research in Neurodegenerative Diseases University of Toronto Toronto Canada

The Centre for Applied Genomics Hospital for Sick Children Toronto Canada

The Mindich Child Health and Development Institute Icahn School of Medicine at Mount Sinai New York NY USA

The Warren Alpert Medical School of Brown University Providence RI USA

Thomas Jefferson University Hospital Philadelphia PA USA

Translational Genomics Research Institute Neurogenomics Division Phoenix AZ USA

Trinity College Dublin Dublin Ireland

UC Gardner Neuroscience Institute Cincinnati OH USA

UCL Genetics Institute University College London London UK

UCL Institute of Neurology London UK

UK Dementia Research Institute Cambridge UK

UK Dementia Research Institute University College London London UK

UMC Utrecht Utrecht The Netherlands

Unidad de Salud Mental Hospital Regional Universitario de Malaga Malaga Spain

Unidad de Trastornos del Movimiento Servicio de Neurología y Neurofisiología Clínica Instituto de Biomedicina de Sevilla Hospital Universitario Virgen del Rocío CSIC Universidad de Sevilla Seville Spain

United Christian Hospital Hong Kong

Universitat Autònoma de Barcelona Barcelona Spain

Université de Lille U1167 RID AGE Risk Factors and Molecular Determinants of Aging Related Diseases Lille France

Université Paul Sabatier Toulouse France

University Clinic of Pediatrics Faculty of Medicine University of Coimbra Coimbra Portugal

University College London London UK

University Health Network University of Toronto Toronto Canada

University Hospital of Wales Cardiff UK

University of Antwerp Antwerp Belgium

University of Bordeaux Bordeaux France

University of California San Francisco CA USA

University of Chicago Chicago IL USA

University of Edinburgh Edinburgh UK

University of Groningen Groningen the Netherlands

University of Lincoln Lincoln UK

University of Liverpool Liverpool UK

University of Montpellier Montpellier France

University of New Mexico Health Sciences Center Albuquerque NM USA

University of Newcastle Callaghan Australia

University of North Carolina at Chapel Hill Chapel Hill NC USA

University of Oslo Oslo Norway

University of Pittsburgh Pittsburgh PA USA

University of Queensland Brisbane Australia

University of Split School of Medicine Split Croatia

University of Toronto Toronto Canada

University of Virginia Center for Public Health Genomics University of Virginia Charlottesville VA USA

UOC Genetica ed Epidemiologica Clinica Az Ospedaliera Padova Italy

VA Boston Healthcare System Boston MA USA

Vall d'Hebron Research Institute Barcelona Spain

Vanderbilt University Medical Center Nashville TN USA

Vilnius University Vilnius Lithuania

Virginia Institute for Psychiatric and Behavioral Genetics Department of Psychiatry Virginia Commonwealth University Richmond VA USA

Walton Centre NHS Foundation Trust Liverpool UK

Washington University School of Medicine St Louis MO USA

Washington University St Louis MO USA

Weill Cornell Medical College New York New York USA

Weill Institute for Neurosciences University of California San Francisco San Francisco CA USA

Wellcome Centre for Human Genetics Nuffield Department of Medicine University of Oxford Oxford UK

Yale Center for Genome Analysis Yale University New Haven CT USA

Yale Child Study Center Yale University School of Medicine New Haven CT USA

Yale University School of Medicine New Haven CT USA

Yong Loo Lin School of Medicine National University of Singapore Singapore

Youthdale Treatment Centers Toronto Canada

Yulius Academy Yulius Mental Health Organization Barendrecht The Netherlands

Zentren für Seelische Gesundheit BBRZ Med Vienna Austria

Zorg op Orde BV Leidschendam The Netherlands