Rhagophthalmidae are a small beetle family known from the eastern Palaearctic and Oriental realms. Rhagophthalmidae are closely related to railroad worms (Phengodidae) and fireflies (Lampyridae) with which they share highly modified paedomorphic females and the ability to emit light. Currently, Rhagophthalmidae include 66 species classified in the following 12 genera: Bicladodrilus Pic, 1921 (two spp.), Bicladum Pic, 1921 (two spp.), Dioptoma Pascoe, 1860 (two spp.), Diplocladon Gorham, 1883 (two spp.), Dodecatoma Westwood, 1849 (eight spp.), Falsophrixothrix Pic, 1937 (six spp.), Haplocladon Gorham, 1883 (two spp.), Menghuoius Kawashima, 2000 (three spp.), Mimoochotyra Pic, 1937 (one sp.), Monodrilus Pic, 1921 (two spp. in two subgenera), Pseudothilmanus Pic, 1918 (two spp.), and Rhagophthalmus Motschulsky, 1854 (34 spp.). The replacement name Haplocladongorhami Kundrata, nom. nov. is proposed for Diplocladonhasseltii Gorham, 1883b (described in subgenus Haplocladon) which is preoccupied by Diplocladonhasseltii Gorham, 1883a. The genus Reductodrilus Pic, 1943 is tentatively placed in Lampyridae: Ototretinae. Lectotypes are designated for Pseudothilmanusalatus Pic, 1918 and P.marginalis Pic, 1918. Interestingly, in the eastern part of their distribution, Rhagophthalmidae have remained within the boundaries of the Sunda Shelf and the Philippines demarcated by the Wallace Line, which separates the Oriental and Australasian realms. This study is intended to be a first step towards a comprehensive revision of the group on both genus and species levels. Additionally, critical problems and prospects for rhagophthalmid research are briefly discussed.

Department of Zoology Faculty of Science Palacky University 17 listopadu 50 77900 Olomouc Czech Republic Palacky University Olomouc Czech Republic

EpiLogic GmbH Agrarbiologische Forschung und Beratung Hohenbachernstr 19 21 85354 Freising Germany EpiLogic GmbH Agrarbiologische Forschung und Beratung Freising Germany

Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P O Box 147100 Gainesville FL 32614 7100 USA Florida Department of Agriculture and Consumer Services Gainesville United States of America

Zobrazit více v PubMed

Amaral DT, Prado RA, Viviani VR. (2012) Luciferase from Fulgeochlizusbruchi (Coleoptera: Elateridae), a Brazilian click-beetle with a single abdominal lantern: molecular evolution, biological function and comparison with other click-beetle luciferases. Photochemical & Photobiological Sciences 11(7): 1259–1267. 10.1039/c2pp25037c PubMed DOI

Amaral DT, Arnoldi FGC, Rosa SP, Viviani VR. (2014) Molecular phylogeny of Neotropical bioluminescent beetles (Coleoptera: Elateroidea) in southern and central Brazil. Luminescence 29(5): 412–422. 10.1002/bio.2561 PubMed DOI

Amaral DT, Mitani Y, Ohmiya Y, Viviani VR. (2016) Organization and comparative analysis of the mitochondrial genomes of bioluminescent Elateroidea (Coleoptera: Polyphaga). Gene 586(2): 254–262. 10.1016/j.gene.2016.04.009 PubMed DOI

Amaral DT, Mitani Y, Oliveira G, Ohmiya Y, Viviani VR. (2017a) Revisiting Coleoptera a1T-rich region: Structural conservation, phylogenetic and phylogeographic approaches in mitochondrial control region of bioluminescent Elateridae species (Coleoptera). Mitochondrial DNA Part A: DNA Mapping, Sequencing, and Analysis 28(5): 671–680. 10.3109/24701394.2016.1174220 PubMed DOI

Amaral DT, Silva JR, Viviani VR. (2017b) Transcriptional comparison of the photogenic and non-photogenic tissues of Phrixothrixhirtus (Coleoptera: Phengodidae) and non-luminescent Chauliognathusflavipes (Coleoptera: Cantharidae) give insights on the origin of lanterns in railroad worms. Gene Reports 7: 78–86. 10.1016/j.genrep.2017.02.004 DOI

Amaral DT, Silva JR, Viviani VR. (2017c) Transcriptomes from the photogenic and non-photogenetic tissues and life stages of the Aspisomalineatum firefly (Coleoptera: Lampyridae): Implications for the evolutionary origins of bioluminescence and its associated light organs. Gene Reports 8: 150–159. 10.1016/j.genrep.2017.07.004 DOI

Amaral DT, Bonatelli IAS, Cerri R, Viviani VR. (2019) Phylogenomic analyses and divergence time estimation of Elateroidea (Coleoptera) based on RNA-Seq data. Comparative Biochemistry and Physiology 30(Part D): 283–289. 10.1016/j.cbd.2019.04.001 PubMed DOI

Arnoldi FGC, Ogoh K, Ohmiya Y, Viviani VR. (2007) Mitochondrial genome sequence of the Brazilian luminescent click beetle Pyrophorusdivergens (Coleoptera: Elateridae): Mitochondrial genes utility to investigate the evolutionary history of Coleoptera and its bioluminescence. Gene 405(1-2): 1–9. 10.1016/j.gene.2007.07.035 PubMed DOI

Ballantyne LA. (1968) Revisional studies of Australian and Indomalayan Luciolini (Coleoptera, Lampyridae, Luciolinae). University of Queensland Papers, Department of Entomology, University of Queensland Press II(6): 103–139.

Ballantyne LA, Lambkin CL. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa 3653(1): 1–162. 10.11646/zootaxa.3653.1.1 PubMed DOI

Ballantyne LA, Lambkin CL, Ho JZ, Jusoh WFA, Nada B, Nak-Eiam S, Thancharoen A, Wattanachaiyingcharoen W, Yiu V. (2019) The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species. Zootaxa 4687(1): 1–174. 10.11646/zootaxa.4687.1.1 PubMed DOI

Bess HA. (1956) Ecological notes on Lamprophorustenebrosus (Walker) (Coleoptera: Lampyridae), an enemy of the giant African snail. Proceedings of the Hawaiian Entomological Society 16: 24–29.

Bezděk J, Regalin R. (2015) Identity of species-group taxa of the Western Palaearctic Clytrini (Coleoptera: Chrysomelidae) described by Maurice Pic and Louis Kocher. Acta Entomologica Musei Nationalis Pragae 55(supplementum): 1–114.

Bi W-X, He J-W, Chen C-C, Kundrata R, Li X-Y. (2019) Sinopyrophorinae, a new subfamily of Elateridae (Coleoptera, Elateroidea) with the first record of a luminous click beetle in Asia and evidence for multiple origins of bioluminescence in Elateridae. ZooKeys 864: 79–97. 10.3897/zookeys.864.26689 PubMed DOI PMC

Blair KG. (1915a) Luminous insects. Nature 96(2406): 411–415. 10.1038/096411a0 DOI

Blair KG. (1915b) Fireflies and glow-worms and their light. The Journal of the Natural History Society of Siam 1: 188–195.

Blair KG. (1915c) Luminous insects. Proceedings of the South London Entomological & Natural History Society 1914 [1915]: 31–45.

Bocak L. (2007) Phengodidae. In: Löbl I, Smetana A. (Eds) Catalogue of Palaearctic Coleoptera, Vol.4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, and Cucujoidea. Apollo Books, Stenstrup, Denmark, 224–225.

Bocak L, Bocakova M, Hunt T, Vogler AP. (2008) Multiple ancient origins of neoteny in Lycidae (Coleoptera): Consequences for ecology and macroevolution. Proceedings of the Royal Society B: Biological Sciences 275(1646): 2015–2023. 10.1098/rspb.2008.0476 PubMed DOI PMC

Bocak L, Barton C, Crampton-Platt A, Chesters D, Ahrens D, Vogler AP. (2014) Building the Coleoptera tree-of-life for >8000 species: Composition of public DNA data and fit with Linnaean classification. Systematic Entomology 39(1): 97–110. 10.1111/syen.12037 DOI

Bocak L, Kundrata R, Andújar Fernández C, Vogler AP. (2016) The discovery of Iberobaeniidae (Coleoptera: Elateroidea), a new family of beetles from Spain, with immatures detected by environmental DNA sequencing. Proceedings of the Royal Society B: Biological Sciences 283(1830): 20152350. 10.1098/rspb.2015.2350 PubMed DOI PMC

Bocak L, Motyka M, Bocek M, Bocakova M. (2018) Incomplete sclerotization and phylogeny: The phylogenetic classification of Plastocerus (Coleoptera: Elateroidea). PLoS ONE 13(3): e0194026. 10.1371/journal.pone.0194026 PubMed DOI PMC

Bocakova M, Bocak L, Hunt T, Teraväinen M, Vogler AP. (2007) Molecular phylogenetics of Elateriformia (Coleoptera): Evolution of bioluminescence and neoteny. Cladistics 23(5): 477–496. 10.1111/j.1096-0031.2007.00164.x DOI

Bogahawatta CNL, Wegiriya HCE, Rajapaksha RSPKM. (2009) Species diversity of Fireflies (Coleoptera: Lampyridae) in selected natural and agricultural habitats in Southern Sri Lanka. Research Journal of the University of Ruhuna, Sri Lanka – Rohana 7: 1–11.

Bouchard P, Bousquet Y, Davies AE, Alonso-Zarazaga MA, Lawrence JF, Lyal CHC, Newton AF, Reid CAM, Schmitt M, Ślipiński SA, Smith ABT. (2011) Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. 10.3897/zookeys.88.807 PubMed DOI PMC

Bourgeois J. (1884) Dascillides et Malacodermes de Nouvelle-Calédonie. Revue d’Entomologie III: 278–290.

Bourgeois J. (1885) Description d'un genre nouveau et d'une espèce nouvelle de Malacodermes de la faune paléarctique. Annales de la Société Entomologique de France 5: 272–274.

Bourgeois J. (1892) Coléoptères du Bengale Occidental. 21e mémoire. Dascillides et Malacodermes. Annales de la Société Entomologique de Belgique 36: 234–238.

Bourgeois J. (1903) Voyage de M. Maurice Maindron dans L'Inde Méridionale (mai à novembre 1901). Dascillides et Malacodermes. Annales de la Société Entomologique de France 72: 478–483.

Bourgeois J. (1905) Voyage du Dr W. Horn a Ceylan. Malacodermes et Lymexylonides. Annales de la Société Entomologique de France 74: 127–133.

Bousquet Y. (2016) Litteratura Coleopterologica (1758–1900): A guide to selected books related to the taxonomy of Coleoptera with publication dates and notes. ZooKeys 583: 1–776. 10.3897/zookeys.583.7084 DOI

Branham MA. (1996) Phengodids: Giant glowworm beetles (A taxonomic survey of lanterns and their use) (continued from Companion 1). Fireflyer Companion 1: 18–19.

Branham MA, Wenzel JW. (2001) The evolution of bioluminescence in cantharoids (Coleoptera: Elateroidea). The Florida Entomologist 84(4): 565–586. 10.2307/3496389 DOI

Branham MA, Wenzel JW. (2003) The origin of photic behavior and the evolution of sexual communication in fireflies (Coleoptera: Lampyridae). Cladistics 19(1): 1–22. 10.1111/j.1096-0031.2003.tb00404.x PubMed DOI

Brues CT. (1941) Characteristics of the larviform female of the Lampyrid beetle, Lamprophorus. Psyche 48(1): 41–44. 10.1155/1941/60591 DOI

Brues CT, Melander AL, Carpenter FM. (1954) Classification of insects. Keys to the living and extinct families of insects, and to the living families of other terrestrial arthropods. Bulletin of the Museum of Comparative Zoology 108: 1–917.

Bugnion E. (1916) Les Insectes phosphorescents. Bulletin de la Murithienne societe Valaisanne des sciences naturelles 39: 82–124.

Bugnion E. (1929) Le ver-luisant provençal et la luciole niçoise. Mémoire Supplément au “Riviera scientifique”. Association des Naturalistes de Nice et des Alpes-Maritimes, Nice, France, 1–131.

Cai C, Tihelka E, Giacomelli M, Lawrence JF, Ślipiński A, Kundrata R, Yamamoto S, Thayer MK, Newton AF, Leschen RAB, Gimmel ML, Lü L, Engel MS, Bouchard P, Huang D, Pisani D, Donoghue PCJ. (2022) Integrated phylogenomics and fossil data illuminate the evolution of beetles. Royal Society Open Science 9(3): 211771. 10.1098/rsos.211771 PubMed DOI PMC

Cardon RP. (1892) Liste des Dascillides et Malacodermes recueillis dans le Chota-Nagpore. Annales de la Société Entomologique de Belgique 36: 238.

Case JF, Strause LG. (1978) 10. Neurally controlled luminescent systems. In: Herring PJ. (Ed.) Bioluminescence in action.Academic Press, London, 331–366.

Chandler DS. (2000) Publication dates of papers on the Anthicidae (Coleoptera) by the Marquis F.T. de LaFerté-Sénectère. Transactions of the American Entomological Society 125 [1999]: 433–439.

Chen TR. (2003) Rhagophthalmidae. In: Chen TR. (Ed.) Fireflies of Taiwan.Field Image Publ., Taipei City, Taiwan, 52–53.

Chen TR, Ho JR. (1996) New record of firefly in Taiwan, Rhagophthalmusohbai Wittmer. Nature Conservation Quarterly 16: 46–49.

Chen TZ, Ho J-Z. (1998) Luminiferous behavior of Rhagophthalmusohbai Wittmer. Nature Conservation Quarterly 23: 34–37. 10.1179/019713698806082967 DOI

Chen Y-F, Yang C-K, Wang C-T, Wang Y-N, Ho J-Z, Yang P-S. (2010) Biology and luminescence behavior of Rhagophthalmusjenniferae (Coleoptera: Rhagophthalmidae) in Xitou Natural Education Area, Experimental Forest, National Taiwan University. Journal of the Experimental Forest of National Taiwan University 24: 195–208. 10.6542/EFNTU.201009_24(3).0005 DOI

Chen X, Dong Z, Liu G, He J, Zhao R, Wang W, Peng Y, Li X. (2019) Phylogenetic analysis provides insights into the evolution of Asian fireflies and adult bioluminescence. Molecular Phylogenetics and Evolution 140: 106600. 10.1016/j.ympev.2019.106600 PubMed DOI

Cicero JM. (1988) Ontophylogenetics of cantharoid larviforms (Coleoptera: Cantharoidea). Coleopterists Bulletin 42: 105–151.

Coblentz WW, Hughes CW. (1926) Spectral energy distribution of the light emitted by plants and animals. Scientific Papers of the Bureau of Standards 21: 521–534. 10.6028/nbsscipaper.228 DOI

Constantin R. (2014) Contribution à la connaissance des Phengodidae de Guyane et description de huit espèces nouvelles (Coleoptera, Elateroidea). Coléoptères de Guyane 8: 86–104.

Constantin R. (2016) Deux nouveaux Phengodidae de Guyane et du Guyana (Coleoptera, Elateroidea). Coleopterist 19: 158–162.

Costa C, Zaragoza-Caballero S. (2010) Phengodidae LeConte, 1861. In: Leschen RAB, Beutel RG, Lawrence JF. (Eds) Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin/New York, 126–135. 10.1515/9783110911213.126 DOI

Costa C, Vanin SA, Casari SA, Viviani VR. (1999) Larvae of neotropical Coleoptera. XXVII. Phrixothrixhirtus: Immatures, neotenic female, adult male and bionomic data (Phengodinae, Phengodidae, Coleoptera). Iheringia, Serie Zoologia, Porto Alegre 86: 9–28.

Crowson RA. (1955) The natural classification of the families of Coleoptera. Nathaniel Lloyd & Co. , Ltd, London, 186 pp.

Crowson RA. (1972) A review of the classification of Cantharoidea (Coleoptera), with the definition of two new families, Cneoglossidae and Omethidae. Revista de la Universidad de Madrid 21: 35–77.

Crowson RA. (1981) The Biology of the Coleoptera. Academic Press, London, 802 pp.

Day JC, Goodall TI, Bailey MJ. (2009) The evolution of the adenylate-forming protein family in beetles: Multiple luciferase gene paralogues in fireflies and glow-worms. Molecular Phylogenetics and Evolution 50(1): 93–101. 10.1016/j.ympev.2008.09.026 PubMed DOI

De Keyzer R. (1989) The cold light of the firefly. Bioluminescence 23: 51–57.

DeCock R. (2004) Larval and adult emission spectra of bioluminescence in three European firefly species. Photochemistry and Photobiology 79(4): 339–342. 10.1562/2003-11-11-RA.1 PubMed DOI

Dong P, Hou Q, Li X, Liang X. (2008) Cloning, expression and sequence analysis of a luciferase gene from the chinese firefly Pyrocoeliapygidialis. Zoological Research 29: 477–484. 10.3724/SP.J.1141.2008.05477 DOI

Dong Z, Yiu V, Liu G, He J, Zhao R, Peng Y, Li X. (2021) Three new species of Lamprigera Motschulsky (Coleoptera, Lampyridae) from China, with notes on known species. Zootaxa 4950(3): 441–468. 10.11646/zootaxa.4950.3.2 PubMed DOI

Douglas HB, Kundrata R, Brunke AJ, Escalona HE, Chapados JT, Eyres J, Richter R, Savard K, Ślipiński A, McKenna D, Dettman JR. (2021) Anchored phylogenomics, evolution and systematics of Elateridae: Are all bioluminescent Elateroidea derived click beetles? Biology 10(6): 451. 10.3390/biology10060451 PubMed DOI PMC

Evenhuis NL. (2020) Catalog of dating sources for zoological works, Fourth edition (completely revised and augmented). Bishop Museum Technical Report 71: 1–369.

Fairmaire L. (1888) Coléoptères de l'Interieur de la Chine. Annales de la Société Entomologique de Belgique 32: 7–46.

Fairmaire L. (1889) Descriptions de Coléoptères de l’Indo-Chine. Annales de la Société Entomologique de France 6 [1888]: 333–378.

Fairmaire L. (1896) Coléoptères de l'Inde et de la Malaisie. Notes from the Leyden Museum 18: 225–238.

Fairmaire L. (1899) Descriptions de Coléoptères nouveaux recueillis en Chine par M. de Latouche. Annales de la Société Entomologique de France 68: 616–643.

Fallon TR, Lower SE, Chang C-H, Bessho-Uehara M, Martin GJ, Bewick AJ, Behringer M, Debat HJ, Wong I, Day JC, Suvorov A, Silva CJ, Stanger-Hall KF, Hall DW, Schmitz RJ, Nelson DR, Lewis SM, Shigenobu S, Bybee SM, Larracuente AM, Oba Y, Weng J-K. (2018) Firefly genomes illuminate parallel origins of bioluminescence in beetles. eLife 7: e36495. 10.7554/eLife.36495 PubMed DOI PMC

Fauvel A. (1904) Faune analytique des Coléoptères de la Nouvelle-Calédonie. 2ème partie. Buprestidae-Tenebrionidae (pars). Revue Entomologie 23: 113–208.

Ferreira VS, Ivie MA. (2022) Lessons from a museum’s cabinet: DNA barcoding and collections-based life stage associations reveals a hidden diversity in the Puerto Rican bank paedomorphic Lycidae (Coleoptera: Elateroidea: Leptolycini). Insect Systematics and Diversity 6(3): 2. 10.1093/isd/ixac006 DOI

Ferreira VS, Keller O, Branham MA, Ivie MA. (2019) Molecular data support the placement of the enigmatic Cheguevaria as a subfamily of Lampyridae (Insecta: Coleoptera). Zoological Journal of the Linnean Society 187(4): 1253–1258. 10.1093/zoolinnean/zlz073 DOI

Ferreira VS, Keller O, Branham MA. (2020) Multilocus phylogeny support the nonbioluminescent firefly Chespirito as a new subfamily in the Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 4(6): 2. 10.1093/isd/ixaa014 DOI

Ferreira VS, Keller O, Ivie MA. (2022) Descriptions of new species of Chespirito Ferreira, Keller & Branham (Coleoptera: Lampyridae: Chespiritoinae) and the first record for the subfamily in the United States. Zootaxa 5124(2): 230–237. 10.11646/zootaxa.5124.2.8 PubMed DOI

Fowler WW. (1912) Coleoptera, general introduction and Cicindelidae and Paussidae. In: Shipley E. (Ed.) Fauna of British India, including Ceylon and Burma.Taylor and Francis, London, 135–138.

Gahan CJ. (1908a) Lampyridae from Ceylon. The Transactions of the Entomological Society of London 1908: xlviii.

Gahan CJ. (1908b) Entomological Society of London Report. Entomologist 41: 205.

Gahan CJ. (1925) Living fireflies from Argentina. In: Green EE (Ed.) The Proceedings of the Entomological Society of London for the year 1924, Wednesday, February 6th, 1924. Transactions of the Entomological Society of London 1925: v–vii.

Ge X, Yuan L, Kang Y, Liu T, Liu H, Yang Y. (2021) Characterization of the first complete mitochondrial genome of Cyphonocerinae (Coleoptera: Lampyridae) with implications for phylogeny and evolution of fireflies. Insects 12(7): 570. 10.3390/insects12070570 PubMed DOI PMC

Ge X, Liu T, Kang Y, Liu H, Yang Y. (2022) First complete mitochondrial genomes of Ototretinae (Coleoptera, Lampyridae) with evolutionary insights into the gene rearrangement. Genomics 114(2): 110305. 10.1016/j.ygeno.2022.110305 PubMed DOI

Geisthardt M, Satô M. (2007) Lampyridae. In: Löbl I, Smetana A. (Eds) Catalogue of Palaearctic Coleoptera, Vol.4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, and Cucujoidea. Apollo Books, Stenstrup, Denmark, 225–234.

Gemminger M. (1869) Tom VI. Rhipidoceridae, Dascillidae, Malacodermidae, Cleridae, Lymexylonidae, Cupesidae, Ptinidae, Bostrychidae, Cioidae. In: Gemminger M, Harold B. (Eds) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus.E. H. Gummi, Monachii, 1609–1800. 10.5962/bhl.title.9089 DOI

Gerstaecker A. (1863) Bericht über die wissenschaftlichen Leistungen im Gebiete der Entomologie während des Jahres 1862. Archiv für Naturgeschichte 29: 315–598.

Goidanich A. (1957) Enciclopedia Agraria Italiana, III. REDA, Roma, Italy, 1168 pp.

Gorham HS. (1880) On the structure of the Lampyridae, with reference to their phosphorescence. The Transactions of the Entomological Society of London 1880: 63–67.

Gorham HS. (1881) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. III. Part 2. Malacodermata. Taylor and Francis, London, xii + 372 pp. [+ 13 pls.]

Gorham HS. (1883a) Description of a new genus and species of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 5–6.

Gorham HS. (1883b) A new subgenus of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 249–250.

Gorham HS. (1883c) Descriptions of Malacodermata in the Civic Museum of Natural History at Genoa. Annali del Museo Civico di Storia Naturale di Genova 18: 591–606.

Gorham HS. (1887) Fam. Malacodermidae. In: Ritsema C. (Ed.) Coleoptera door verschillende specialiteiten bewerkt en tot een geheel bijeengebracht.In: Snelleman JF (Ed.) Bijdragem tot de Kennis der Fauna van Midden-Sumatra. Eerste Deel. EJ Brill, Leiden, 60–77.

Gorham HS. (1890) XVI. Notes on the species of the families Lycidae and Lampyridae, contained in the Imperial Museum of Calcutta, with descriptions of new species, and a list of the species at present described from India. The Transactions of the Entomological Society of London 1890: 541–550. 10.1111/j.1365-2311.1890.tb02707.x DOI

Gorham HS. (1895) List of the Coleoptera in the collection of H. E. Andrewes, Esq., from India and Burma, with descriptions of new species and notes. Families: Malacodermata, Erotylidae, Endomychidae. Annales de la Société Entomologique de Belgique 39: 293–330.

Gorham HS. (1903) On Coleoptera collected in India by MM. H.-E. and H.-L. Andrewes. Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae. Annales de la Société Entomologique de Belgique 47: 323–330.

Goto Y, Kawashima I. (2000) Distributional records of some fireflies (Coleoptera, Lampyridae and Rhagophthalmidae) from the Ryukyu Islands, Southwest Japan. Elytra, Tokyo 28: 141–144.

Gravely FH. (1915) Notes on the habits of Indian insects, myriapods and arachnids. Records of the Indian Museum 11: 483–539. 10.26515/rzsi/v11/i6/1915/163052 DOI

Green EE. (1913) On some luminous Coleoptera from Ceylon. Transactions of the Entomological Society of London 1912/1913 [1912]: 717–719. 10.1111/j.1365-2311.1913.tb03114.x DOI

Hall R. (1998) The plate tectonics of Cenozoic SE Asia and the distribution of land and sea. In: Hall R, Holloway JD. (Eds) Biogeography and Geological Evolution of SE Asia.Backhuys Publishers, Leiden, 99–131.

Halverson RC, Case JF, Buck JB, Tiemann DL. (1973) Control of luminescence in phengodid beetles. Journal of Insect Physiology 19(6): 1327–1339. 10.1016/0022-1910(73)90215-1 DOI

Handlirsch A. (1925) Systematische Übersicht. In: Schröder C. (Ed.) Handbuch der Entomologie.Band III. Geschichte, Literatur, Technik, Palaontologie, Phylogenie, Systematik. Verlag von Gustav Fischer, Jena, 377–1139.

Haneda Y. (1950) Star-worm in Singapore. Shin Konchu 3: 2–5. [New Insect]

Haneda Y. (1955) Luminous organisms of Japan and the Far East. In: Johnson FH. (Ed.) The luminescense of biological systems.AAAS, Washington DC, 335–385.

Haneda Y. (1985) Luminous organisms. Kōseisha Kōseikaku, Tokyo, Japan, 318 pp.

Harvey EN. (1952) Coleoptera. In: Harvey EN. (Ed.) Bioluminescence.Academic Press, New York, 389–471.

Harvey EN. (1955) Survey of luminous organisms: Problems and prospects. In: Johnson FH. (Ed.) The luminescense of biological systems.AAAS, Washington DC, 1–24.

Harvey EN. (1957) A History of Luminescence from the Earliest Times Until 1900. The American Philosophical Society, Independence Square, Philadelphia, 692 pp. 10.5962/bhl.title.14249 DOI

Hayashi F, Suzuki H. (2003) Fireflies with or without prespermatophores: Evolutionary origins and life-history consequences. Entomological Science 6(1): 3–10. 10.1046/j.1343-8786.2003.00003.x DOI

He J, Bi W, Dong Z, Liu G, Zhao R, Wang W, Li X. (2019) The mitochondrial genome of the first luminous click-beetle (Coleoptera: Elateridae) recorded in Asia. Mitochondrial DNA Part B 4(1): 565–567. 10.1080/23802359.2018.1555019 DOI

He J, Yao Y, Dong Z, Ruan Y, Chang Z, Zhao R, Wang W, Li X. (2022) Complete mitochondrial genome of Pectocera sp. (Elateridae: Dendrometrinae: Oxynopterini) and its phylogenetic implications. Archives of Insect Biochemistry and Physiology 111(1): e21957. 10.1002/arch.21957 PubMed DOI

Herring PJ. (1978) Appendix: A classification of Luminous Organisms. In: Herring PJ. (Ed.) Bioluminescence in action.Academic Press, London, 461–476.

Herring PJ. (1987) Systematic distribution of bioluminescence in living organisms. Journal of Bioluminescence and Chemiluminescence 1(3): 147–163. 10.1002/bio.1170010303 PubMed DOI

Heyden L. (1886) Beiträge zur Coleopteren-Fauna von Pecking in Nord-China. Deutsche Entomologische Zeitschrift 30: 281–292. 10.1002/mmnd.48018860216 DOI

Ho J-Z. (2002) Larval morphology of twenty one species and bionomics of fireflies (Coleoptera: Lampyridae) in Taiwan. Ph.D. Thesis, National Chung Hsing University, Taichung, 162 pp.

Ho J-Z, Chen Y-F, Cheng S-H, Tsai X-L, Yang P-S. (2012) Two new species of Rhagophthalmus Motschulsky (Coleoptera: Rhagophthalmidae) from Matzu Archipelago, Taiwan with biological commentary. Zootaxa 3274(1): 1–13. 10.11646/zootaxa.3274.1.1 DOI

Hoffmann KH. (1984) Environmental aspects of insect bioluminescence. In: Hoffmann KH. (Ed.) Environmental Physiology and Biochemistry of Insects.Springer-Verlag, Berlin, Heidelberg, 225–244. 10.1007/978-3-642-70020-0_9 DOI

Hosoe T, Saito K, Ichikawa M, Ohba N. (2014) Chemical defense in the firefly, Rhagophthalmusohbai (Coleoptera: Rhagophthalmidae). Applied Entomology and Zoology 49(2): 331–335. 10.1007/s13355-014-0255-1 DOI

Hua LZ. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp.

Hunt T, Bergsten J, Levkanicova Z, Papadopoulou A, John OS, Wild R, Hammond PM, Ahrens D, Balke M, Caterino MS, Gómez-Zurita J, Ribera I, Barraclough TG, Bocakova M, Bocak L, Vogler AP. (2007) A comprehensive phylogeny of beetles reveals the evolutionary origins of a superradiation. Science 318(5858): 1913–1916. 10.1126/science.1146954 PubMed DOI

ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, 305 pp.

ICZN [International Commission on Zoological Nomenclature] (2014) Opinion 2340 (Case 3592) Dodecatoma Westwood, 1849 (Insecta, Coleoptera): name conserved by suppression of Dodecatoma Dufour, 1841 (Insecta, Plecoptera). The Bulletin of Zoological Nomenclature 71: 195–196. 10.21805/bzn.v71i3.a4 DOI

Jakobson GG. (1911) Zhuki Rossii i zapadnoy Evropy. Fasc. 9. St. Petersburg, Izdaniye A. Devriena, 653–687.

Janisova K, Bocakova M. (2013) Revision of the subfamily Ototretinae (Coleoptera: Lampyridae). Zoologischer Anzeiger 252(1): 1–19. 10.1016/j.jcz.2012.01.001 DOI

Jeng M-L. (2008) Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera). PhD Thesis, University of Kansas, Lawrence, Kansas, USA, 388 pp.

Jeng M-L. (2019) Biofluorescence in terrestrial animals, with emphasis on fireflies: A review and field observation. In: Suzuki H. (Ed.) Bioluminescence – Analytical Applications and Basic Biology.IntechOpen, London, 1–25. 10.5772/intechopen.86029 DOI

Jeng M-L, Lai J, Yang P-S, Satô M. (2000) Notes on the taxonomy of Lamprigerayunnana (Fairmaire) and the genus Lamprigera Motschulsky (ColeopteraLampyridae). Japanese Journal of Systematic Entomology 6: 313–319.

Johnson PJ, Dewalt E, Evenhuis N. (2012) Case 3592 Dodecatoma Westwood, 1849 (Insecta, Coleoptera): Proposed conservation by suppression of Dodecatoma Dufour, 1841 (Insecta, Plecoptera). Bulletin of Zoological Nomenclature 69(3): 178–181. 10.21805/bzn.v69i3.a10 DOI

Junk W. (1912) Bibliographia coleopterologica. W. Junk, Berlin, 1–146. 10.1007/978-94-017-6615-9_1 DOI

Kawashima I. (1998) Morphology of larviform adult females in Japanese Cantharoidea, with special reference to two families and genera Rhagophthalmus (Rhagophthalmidae) and Stenocladius (Lampyridae). The Nature and Insects 33: 16–18.

Kawashima I. (1999) The lampyrid beetles of the genus Stenocladius (Coleoptera, Lampyridae) of the Ryukyu Islands, Southwest Japan with descriptions of two new species. Elytra, Tokyo 27: l41–l158.

Kawashima I. (2000) A redescription of Rhagophthalmusingens Fairmaire (Coleoptera, Rhagophthalmidae) from northern Vietnam, with establishment of a new genus in the family. Elytra, Tokyo 28: 131–140.

Kawashima I. (2002) A new species of the Rhagophthalmid genus Menghuoius (Coleoptera, Rhagophthalmidae) from Myanmar. Elytra, Tokyo 30: 487–492.

Kawashima I, Satô M. (2001) Three new species of the genus Rhagophthalmus (Coleoptera, Rhagophthalmidae) from southeast Asia. Elytra, Tokyo 29: 423–434.

Kawashima I, Sugaya H. (2003) An additional new species of the genus Rhagophthalmus (Coleoptera, Rhagophthalmidae) from Taiwan, with a key to the males of the Taiwanese and Japanese species. Elytra, Tokyo 31: 353–359.

Kawashima I, Suzuki H, Satô M. (2003) A check-list of Japanese fireflies (Coleoptera, Lampyridae and Rhagophthalmidae). Japanese Journal of Systematic Entomology 9: 241–261.

Kawashima I, Lawrence JF, Branham MA. (2010) Rhagophthalmidae Olivier, 1907. In: Leschen RAB, Beutel RG, Lawrence JF. (Eds) Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin/New York, 135–140. 10.1515/9783110911213.135 DOI

Kazantsev SV. (2012) Two new species of Rhagophthalmidae (Insecta: Coleoptera) from the Himalayas. In: Hartmann M, Weipert J. (Eds) Biodiversity and Natural Heritage of the Himalaya IV.Verein der Freunde und Förderer des Naturkundemuseums Erfurt e. V., Erfurt, Germany, 349–352.

Kim I, Lee SC, Bae JS, Jin BR, Kim SE, Kim JK, Yoon HJ, Yang SR, Lim SH, Sohn HD. (2000) Genetic divergence and phylogenetic relationships among the Korean fireflies, Hotariapapariensis, Luciolalateralis, and Pyrocoeliarufa (Coleoptera: Lampyridae), using mitochondrial DNA sequences. Korean Journal of Applied Entomology 39: 211–226.

Kobayashi Y, Suzuki H, Ohba N. (2001) Formation of a spherical germ rudiment in the glow-worm, Rhagophthalmusohbai Wittmer (Coleoptera: Rhagophthalmidae), and its phylogenetic implications. Proceedings of the Arthropodan Embryological Society of Japan 36: 1–5.

Kobayashi Y, Suzuki H, Ohba N. (2002) Embryogenesis of the glowworm Rhagophthalmusohbai Wittmer (Insecta: Coleoptera, Rhagophthalmidae), with emphasis on the germ rudiment formation. Journal of Morphology 253(1): 1–9. 10.1002/jmor.1109 PubMed DOI

Kobayashi Y, Suzuki H, Ohba N. (2003) Development of pleuropodia in the embryo of the glowworm Rhagophthalmusohbai (Rhagophthalmidae, Coleoptera, Insecta), with comments on their probable function. Proceedings of the Arthropodan Embryological Society of Japan 38: 19–26.

Kovalev AV, Kirejtshuk AG. (2016) Asiopsectra gen. n., a second genus of the family Brachypsectridae (Coleoptera, Elateroidea) from the Palaearctic Region. Insect Systematics & Evolution 47(3): 195–208. 10.1163/1876312X-47022140 DOI

Kundrata R, Bocak L. (2011a) Redescription and relationships of Pseudothilmanus Pic (Coleoptera: Rhagophthalmidae)—a long-term neglected glow-worm beetle genus from the Himalayas. Zootaxa 2794(1): 57–62. 10.11646/zootaxa.2794.1.4 DOI

Kundrata R, Bocak L. (2011b) The phylogeny and limits of Elateridae (Insecta, Coleoptera): Is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40(4): 364–378. 10.1111/j.1463-6409.2011.00476.x DOI

Kundrata R, Bocak L. (2019) Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies Sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187(2): 413–452. 10.1093/zoolinnean/zlz033 DOI

Kundrata R, Bocakova M, Bocak L. (2013) The phylogenetic position of Artematopodidae (Coleoptera: Elateroidea), with description of the first two Eurypogon species from China. Contributions to Zoology 82(4): 199–208. 10.1163/18759866-08204004 DOI

Kundrata R, Bocakova M, Bocak L. (2014) The comprehensive phylogeny of the superfamily Elateroidea (Coleoptera: Elateriformia). Molecular Phylogenetics and Evolution 76: 162–171. 10.1016/j.ympev.2014.03.012 PubMed DOI

Kundrata R, Gunter NL, Douglas H, Bocak L. (2016) Next step toward a molecular phylogeny of click-beetles (Coleoptera: Elateridae): redefinition of Pityobiinae, with a description of a new subfamily, Parablacinae, from the Australasian Region. Austral Entomology 55(3): 291–302. 10.1111/aen.12185 DOI

Kundrata R, Jäch MA, Bocak L. (2017) Molecular phylogeny of the Byrrhoidea-Buprestoidea complex (Coleoptera, Elateriformia). Zoologica Scripta 46(2): 150–164. 10.1111/zsc.12196 DOI

Kundrata R, Blank SM, Prosvirov AS, Sormova E, Gimmel ML, Vondráček D, Kramp K. (2019) One less mystery in Coleoptera systematics: The position of Cydistinae (Elateriformiaincertae sedis) resolved by multigene phylogenetic analysis. Zoological Journal of the Linnean Society 187(4): 1259–1277. 10.1093/zoolinnean/zlz104 DOI

Kusy D, Motyka M, Andujar C, Bocek M, Masek M, Sklenarova K, Kokas F, Bocakova M, Vogler AP, Bocak L. (2018a) Genome sequencing of Rhinorhipus Lawrence exposes an early branch of the Coleoptera. Frontiers in Zoology 15(1): 21. 10.1186/s12983-018-0262-0 PubMed DOI PMC

Kusy D, Motyka M, Bocek M, Vogler AP, Bocak L. (2018b) Genome sequences identify three families of Coleoptera as morphologically derived click beetles (Elateridae). Scientific Reports 8(1): 17084. 10.1038/s41598-018-35328-0 PubMed DOI PMC

Kusy D, He J-W, Bybee SM, Motyka M, Bi W-X, Podsiadlowski L, Li X-Y, Bocak L. (2021) Phylogenomic relationships of bioluminescent elateroids define the ‘lampyroid’ clade with clicking Sinopyrophoridae as its earliest member. Systematic Entomology 46(1): 111–123. 10.1111/syen.12451 DOI

Lacordaire MT. (1857) Histoire naturelle des insectes. Genera des Coléoptères. Tome Quatrième. Librairie Encyclopédique de Roret, Paris, 579 pp.

Lau TFS, Meyer-Rochow VB. (2006) Sexual dimorphism in the compound eye of Rhagophthalmusohbai (Coleoptera: Rhagophthalmidae): I. Morphology and ultrastructure. Journal of Asia-Pacific Entomology 9(1): 19–30. 10.1016/S1226-8615(08)60271-X DOI

Lau TFS, Ohba N, Arikawa K, Meyer-Rochow VB. (2007) Sexual dimorphism in the compound eye of Rhagophthalmusohbai (Coleoptera: Rhagophthalmidae): II. Physiology and function of the eye of the male. Journal of Asia-Pacific Entomology 10(1): 27–31. 10.1016/S1226-8615(08)60327-1 DOI

Lawrence JF. (1982) Coleoptera. In: Parker SP. (Ed.) Synopsis and classification of living organisms.McGraw-Hill Inc., New York, 483–553.

Lawrence JF. (1988) Rhinorhiphidae, a new beetle family from Australia, with comments on the phylogeny of the Elaterformia. Invertebrate Taxonomy 2(1): 1–53. 10.1071/IT9880001 DOI

Lawrence JF. (2016) Classification (families & subfamilies). In: Beutel RG, Leschen RAB. (Eds) Coleoptera, Beetles, Vol.1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim), 2nd Edn. In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH: Berlin/Boston, 13–22. 10.1515/9783110373929-005 DOI

Lawrence JF, Newton AF. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In: Pakaluk J, Ślipiński A. (Eds) Biology, Phylogeny, and Classification of Coleoptera.Papers Celebrating the 80th Birthday of Roy A Crowson, vol 1, 2. Muzeum i Instytut Zoologii PAN, Warszawa, 779–1083.

Lawrence JF, Beutel RG, Leschen RAB, Ślipiński A. (2010a) Changes in classification and list of families and subfamilies. In: Leschen RAB, Beutel RG, Lawrence JF. (Eds) Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin/New York, 1–7. 10.1515/9783110911213.1 DOI

Lawrence JF, Kawashima I, Branham MA. (2010b) ElateriformiaIncertae Sedis. In: Leschen RAB, Beutel RG, Lawrence JF. (Eds) Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin/New York, 162–177. 10.1515/9783110911213.162 DOI

Lawrence JF, Ślipiński SA, Seago AE, Thayer MK, Newton AF, Marvaldi AE. (2011) Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annales Zoologici 61(1): 1–217. 10.3161/000345411X576725 DOI

Lawrence JF, Zhou Y-L, Lemann C, Sinclair B, Ślipiński A. (2021) The hind wing of Coleoptera (Insecta): Morphology, nomenclature and phylogenetic significance. Part 1. General discussion and Archostemata-Elateroidea. Annales Zoologici 71(3): 421–606. 10.3161/00034541ANZ2021.71.3.001 DOI

Lefroy HM. (1909) Indian Insect Life. A manual of the insects of the plains (tropical India). Calcutta & Simla, Thacker, Spink & Co., W. Thacker & Co. , 2 Creed Lane, London, 786 pp.

LeSage L. (1991) Phengodidae (Cantharoidea) (including Rhagophthalmidae). Lawrence JF (Ed.) Order Coleoptera. In: Stehr FW. (Ed.) Immature Insects.Vol. 2. Kendall/Hunt Publishing Company, Dubuque, Iowa, 424–426.

Levkanicova Z, Bocak L. (2009) Identification of net-winged beetle larvae (Coleoptera: Lycidae) using three mtDNA fragments: a comparison of their utility. Systematic Entomology 34(2): 210–221. 10.1111/j.1365-3113.2008.00457.x DOI

Li X, Liang X. (2008) A gigantic bioluminescent starworm (Coleoptera: Rhagophthalmidae) from Northwest Yunnan, China. Entomological News 119(2): 109–112. 10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2 DOI

Li X, Yang S, Xie M, Liang X. (2006) Phylogeny of fireflies (Coleoptera: Lampyridae) inferred from mitochondrial 16S ribosomal DNA, with references to morphological and ethological traits. Progress in Natural Science 16(8): 817–826. 10.1080/10020070612330074 DOI

Li X, Ogoh K, Ohba N, Liang X, Ohmiya Y. (2007) Mitochondrial genomes of two luminous beetles, Rhagophthalmuslufengensis and R.ohbai (Arthropoda, Insecta, Coleoptera). Gene 392(1-2): 196–205. 10.1016/j.gene.2006.12.017 PubMed DOI

Li X, Ohba N, Liang X. (2008a) Two new species of Rhagophthalmus Motschulsky (Coleoptera: Rhagophthalmidae) from Yunnan, south-western China, with notes on known species. Entomological Science 11(2): 259–267. 10.1111/j.1479-8298.2008.00264.x DOI

Li X, Xie M, Liang X. (2008b) Rhagophthalmid – a special group of fireflies. Chinese Bulletin of Entomology 45: 494–498.

Li K-Q, Wang Y-Z, Dong D-Z, Zhang L-K. (2015) Catalog of insect type specimens preserved at the Kunming Institute of Zoology, Chinese Academy of Science with corrections of some specimens. Zoological Research 36: 263–284. PubMed PMC

Li Y-D, Kundrata R, Packova G, Huang D, Cai C. (2021a) A unique elateroid lineage from mid-Cretaceous Burmese amber (Coleoptera: Elateroidea). Scientific Reports 11(1): 21985. 10.1038/s41598-021-01398-w PubMed DOI PMC

Li Y-D, Kundrata R, Tihelka E, Liu Z, Huang D, Cai C. (2021b) Cretophengodidae, a new Cretaceous beetle family, sheds light on the evolution of bioluminescence. Proceedings of the Royal Society B 288(1943): 20202730. 10.1098/rspb.2020.2730 PubMed DOI PMC

Liu H-Y, Kang Z-X, Zhang F, Ge X-Y, Yang Y-X. (2019) The complete mitogenome of Lycostomus sp. (Elateroidea: Lycidae). Mitochondrial DNA Part B 4(2): 3813–3815. 10.1080/23802359.2019.1682483 PubMed DOI PMC

Liu G-C, Dong Z-W, Hou Q-B, He J-W, Zhao R-P, Wang W, Li X-Y. (2020) Second rhagophthalmid luciferase cloned from Chinese glow-worm Menghuoiusgiganteus (Rhagophthalmidae: Elateroidea). Photochemistry and Photobiology 96(1): 46–54. 10.1111/php.13172 PubMed DOI

Lloyd JE. (1971) Bioluminescent communication in insects. Annual Review of Entomology 16(1): 97–122. 10.1146/annurev.en.16.010171.000525 DOI

Lloyd JE. (1978) Insect Bioluminescence. Chap. 8. In: Herring PJ. (Ed.) Bioluminescence in action.Academic Press, London, 241–272.

Lloyd JE. (1979) Sexual selection in luminescent beetles. In: Blum MS, Blum NA. (Eds) Sexual selection and reproductive competition in insects.Academic Press, New York, NY, 293–342. 10.1016/B978-0-12-108750-0.50015-X DOI

Lloyd JE. (1983) Bioluminescence and communication in insects. Annual Review of Entomology 28(1): 131–160. 10.1146/annurev.en.28.010183.001023 DOI

Lohman DJ, de Bruyn M, Page T, von Rintelen K, Hall R, Ng PKL, Shi H-T, Carvalho GP, von Rintelen T. (2011) Biogeography of the Indo-Australian archipelago. Annual Review of Ecology, Evolution, and Systematics 42(1): 205–226. 10.1146/annurev-ecolsys-102710-145001 DOI

Lomolino MV, Riddle BR, Whittaker RJ. (2017) Biogeography: biological diversity across space and time. 5th Edn. Sinauer Associates Inc. , Sunderland, Massachusetts, 759 pp.

Lucas R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius. Pars 1. Nicolaische Verlags-Buchhandlung R. Stricker, Berlin, 696 pp.

Marschall AF. (1873) Nomenclator zoologicus continens nomina systematica generum animalium tam viventium quam fossilium, secundum ordinem alphabeticum disposita, sub auspiciis et sumptibus C.R. Societatis zoologico-botanicae conscriptus a comite Augusto de Marschall, Typis Caroli Ueberreuter (M. Salzer), Vindobonae, 482 pp. 10.5962/bhl.title.1216 DOI

Martin GJ, Lord NP, Branham MA, Bybee SM. (2015) Review of the firefly visual system (Coleoptera: Lampyridae) and evolution of the opsin genes underlying color vision. Organisms, Diversity & Evolution 15(3): 513–526. 10.1007/s13127-015-0212-z DOI

Martin GJ, Branham MA, Whiting MF, Bybee SM. (2017) Total evidence phylogeny and the evolution of adult bioluminescence in fireflies (Coleoptera: Lampyridae). Molecular Phylogenetics and Evolution 107: 564–575. 10.1016/j.ympev.2016.12.017 PubMed DOI

Martin GJ, Stanger-Hall KF, Branham MA, Silveira LFL, Lower SE, Hall DW, Li X-Y, Lemmon AR, Lemmon EM, Bybee SM. (2019) Higher-Level Phylogeny and Reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3(6): 11. 10.1093/isd/ixz024 DOI

McDermott FA. (1914) The ecologic relations of the photogenic function among insects. Zeitschrift für wissenschaftliche Insektenbiologie 10: 303–307. 10.1007/BF02087204 DOI

McDermott FA. (1964) The taxonomy of the Lampyridae (Coleoptera). Transactions of the American Entomological Society 90: 1–72.

McDermott FA. (1966) Lampyridae. In: Steel WO. (Ed.) Coleopterorum Catalogus Supplementa, Pars 9.Uitgeverij Dr. W. Junk, Gravenhage, 1–149.

McElroy WD, Seliger HH, De Luca M. (1974) Insect Bioluminescence. In: Rockstein M. (Ed.) The Physiology of Insecta, 2nd Edn., Vol. II, Academic Press, NY, USA, 411–460. 10.1016/B978-0-12-591602-8.50015-1 DOI

McKenna DD, Wild AL, Kanda K, Bellamy CL, Beutel RG, Caterino MS, Farnum CW, Hawks DC, Ivie MA, Jameson ML, Leschen RAB, Marvaldi AE, McHugh JV, Newton AF, Robertson JA, Thayer MK, Whiting MF, Lawrence JF, Ślipiński A, Maddison DR, Farrell BD. (2015) The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40(4): 835–880. 10.1111/syen.12132 DOI

McKenna DD, Shin S, Ahrens D, Balke M, Beza-Beza C, Clarke DJ, Donath A, Escalona HE, Friedrich F, Letsch H, Liu S, Maddison D, Mayer C, Misof B, Murin PJ, Niehuis O, Peters RS, Podsiadlowski L, Pohl H, Scully ED, Yan EV, Zhou X, Ślipiński A, Beutel RG. (2019) The evolution and genomic basis of beetle diversity. Proceedings of the National Academy of Sciences of the United States of America 116(49): 24729–24737. 10.1073/pnas.1909655116 PubMed DOI PMC

Mikšić R, Mikšić S. (1966) Lampyridae Jugoslavije. Akademija nauka i umjetnosti Bosne i Hercegovine 29: 29–68.

Moreira AC, Amaral DT, Gabriel GVM, Viviani VR. (2022) Cloning and molecular properties of a novel luciferase from the Brazilian Bicellonychalividipennis (Lampyridae: Photurinae) firefly: comparison with other firefly luciferases. Photochemical & Photobiological Sciences 21(9): 1559–1571. 10.1007/s43630-022-00240-0 PubMed DOI

Morice FD. (1913) The Proceedings of the Entomological Society of London for the year 1912, Wednesday, November 6th, 1912. Transactions of the Entomological Society of London 1912/1913 [1912]: civ–cxviii.

Motschulsky V. (1854) Diagnoses de Coléoptères nouveaux, trouvés par M. M. Tararinoff et Guschkéwitsch aux environs de Pékin. Études entomologiques 2: 44–51.

Motschulsky V. (1859) Insectes des Indes orientales, et de contrées analogues. Études entomologiques 8: 25–146.

Motschulsky V. (1861) Essai d’un Catalogue des Insectes de l’ile Ceylan. Bulletin de la Société Impériale des Naturalistes de Moscou 34: 95–155.

Nakane T. (1968) The taxonomy of Lampyridae. Konchu to Shizen 3: 3–6. [in Japanese] [Insects and Nature]

Nakane T. (1997) On the species of the genus Stenocladius in Japan. Konchu to Shizen 32: 33–36. [Insects and Nature]

Noguchi T, Ikeda M, Ohmiya Y, Nakajima Y. (2008) Simultaneous monitoring of independent gene expression patterns in two types of cocultured fibroblasts with different color-emitting luciferases. BMC Biotechnology 8(1): 40. 10.1186/1472-6750-8-40 PubMed DOI PMC

O’Keefe (2002) Family 61. Phengodidae. In: Arnett Jr RH, Thomas MC, Skelley PE, Frank JH. (Eds) American Beetles, Vol.2., Polyphaga: Scarabaeoidea through Curculionoidea. CRC, Boca Raton, FL, 181–186.

Oba Y. (2015) Insect bioluminescence in the post-molecular biology era. In: Hoffmann KK. (Ed.) Insect Molecular Biology and Ecology.CRC Press: Boca Raton, Florida, 94–120.

Oba Y, Branham MA, Fukatsu T. (2011) The terrestrial bioluminescent animals of Japan. Zoological Science 28(11): 771–789. 10.2108/zsj.28.771 PubMed DOI

Ohba N. (1980) Bioluminescent animals of the land. Animals and Nature 10: 13–20.

Ohba N. (1995) Synchronous flashing of Pterotyxcribellata fireflies and mimicry by other insects. Konchu to Shizen 30: 9–14. [Insects and Nature]

Ohba N. (1997a) Twenty years with fireflies – an outline of research in Japan. The Insectarium 34: 4–18.

Ohba N. (1997b) Group flashing fire-flies and fire-fly mimicry in tropical Asia. Biological Sciences 49: 17–22.

Ohba N. (1997c) Breeding of the firefly, Rhagophthalmusohbai (Coleoptera: Rhagophthalmidae). Science Report of the Yokosuka City Museum 45: 51–55.

Ohba N. (1998) Biology of the fireflies. Konchu to Shizen 33: 2–6. [Insects and Nature]

Ohba N. (2004a) Flash communication systems of Japanese fireflies. Integrative and Comparative Biology 44(3): 225–233. 10.1093/icb/44.3.225 PubMed DOI

Ohba N. (2004b) Fireflies study as integrative science. Konchu to Shizen 39: 4–8. [Insects and Nature]

Ohba N, Goto Y, Kawashima I. (1996a) External morphology and behavior of Rhagophthalmusohbai Wittmer, 1994 (Coleoptera; Rhagophthalmidae) and its habitat. Science Report of the Yokosuka City Museum 44: 1–19.

Ohba N, Goto Y, Kawashima I. (1996b) External morphology, color-marking patterns and habits of the larval stage in genus Stenocladius (Coleoptera: Lampyridae). Science Report of the Yokosuka City Museum 44: 21–31.

Ohba N, Goto Y, Kawashima I. (1997) Behavior and adult female morphology of firefly, genus Stenocladius (Coleoptera: Lampyridae) in Japan. Science Report of the Yokosuka City Museum 45: 23–37.

Ohmiya Y, Sumiya M, Viviani VR, Ohba N. (2000) Comparative aspects of a luciferase molecule from the Japanese luminous beetle, Rhagophthalmusohbai. Science Report of the Yokosuka City Museum 47: 31–38.

Olivier E. (1885) Catalogue des Lampyrides faisant partie des collections du Musée civique de Gênes. Annali del Museo Civico di Storia Naturale di Genova Serie 2 22: 333–374.

Olivier E. (1902) Catalogue des espèces de Luciola et genres voisins decrits jusqu’a ce jour. Revue Scientifique du Bourbonnais et du Centre de la France 15: 69–88.

Olivier E. (1907) 53me Fascicule. Coleoptera. Fam. Lampyridae. In: Genera Insectorum. Vol. VIII. Verteneuil & Desmet, Bruxelles, Belgium, 1–74.

Olivier E. (1908) Communications. Description d’un nouveau genre de Lampyrides. Bulletin de la Société Entomologique de France 1908: 17–18. 10.3406/bsef.1908.24253 DOI

Olivier E. (1910) Rhagophthalmidae, Drilidae. In: Junk W, Schenkling S. (Eds) Coleopterorum Catalogus, pars 10.W. Junk, Berlin, Germany, 1–10. 10.1007/978-94-011-9697-0_1 DOI

Olivier E. (1911) Lampyrides et Drilide nouveaux de la collection de M. Veth. Tijdschrift voor Entomologie 54: 16–20. 10.3406/bsef.1911.24878 DOI

Olivier E. (1912) Revision du genre Rhagophthalmus (Col. Lampyr.) et descriptions d’espèces nouvelles. Annales de la Société entomologique de France 80 [1911]: 467–472.

Pascoe FP. (1860) Notices of new or little known genera and species of Coleoptera. Part II. Journal of Entomology 1: 98–132. [plates V–VIII.]

Pascoe FP. (1862) Notices of new or little known genera and species of Coleoptera. Part III. Journal of Entomology 1: 319–370.

Paulus HF. (1972) Die systematische und phylogenetische Stellung der Karumiidae, mit einer beschreibung von Escalerinaserraticornis n.sp. aus S-Persien. Senckenbergiana Biologica 53: 37–54.

Paulus HF. (1975) Penicillophorusctenotarsus n. gen. et n. sp. aus Kolumbien, mit einer Beschreibung einer neuen Tribus Penicillophorini der Phengodidae (Col., Polyphaga, Cantharoidea). Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 25(1973/1974): 69–80.

Pic M. (1914) Nouveaux Coléoptères de diverses familles. Mélanges Exotico-Entomologiques 10: 7–20.

Pic M. (1916) Diagnoses génériques et spécifiques. Mélanges Exotico-Entomologiques 18: 2–20.

Pic M. (1917) Descriptions abrégées diverses. Mélanges Exotico-Entomologiques 23: 2–20.

Pic M. (1918) Courtes descriptions diverse. Mélanges Exotico-Entomologiques 28: 1–24.

Pic M. (1921a) Diagnoses de Coléoptères exotiques. L'Échange. Revue Linnéenne 37: 15–16.

Pic M. (1921b) Nouveautés diverses. Mélanges Exotico-Entomologiques 33: 1–32.

Pic M. (1923) Étude des Malacodermes de L'Indochine recueillis par M. R. Vitalis de Salvaza. Faune Entomologique de l'Indochine Française 6: 7–63.

Pic M. (1924) Lampyrides et Telephorides des iles Philippines. Philippine Journal of Science 25: 711–731.

Pic M. (1925a) Malacodermes exotiques. L'Échange. Revue Linnéenne 41: 17–20.

Pic M. (1925b) Nouveaux Malacodermes Asiatiques. Bulletin du Muséum National d'Histoire Naturelle 31: 72–73.

Pic M. (1928) Un nouveau Drilide (Col). Bulletin bi-mensuel de la Société Linnéenne de Lyon 7: 86–87.

Pic M. (1930a) Dr. E. Mjöberg's Zoological Collections from Sumatra. 10. Dascillidae et Malacodermata. Arkiv för Zoologi 21: 1–6.

Pic M. (1930b) Contribution a L'Étude des Coléoptères Malacodermes (2e article). Annales de la Société Entomologique de France 99: 311–324.

Pic M. (1937) Malacodermes exotiques. L'Échange. Revue Linnéenne 53: 137–148.

Pic M. (1938) Malacodermata. In: Sjöstedt Y. (Ed.) Insekten aus China im Naturhistorischen Reichsmuseum zu Stockholm.Heimgebracht von Direktor Kjell Kolthoff und anderen Schwedischen Forschern und Reisende. Arkiv för Zoologi 30A, 15–16.

Pic M. (1943) Coléoptères du globe (suite). L'Échange. Revue Linnéenne 59: 9–12.

Pic M. (1951) Notes et descriptions. Diversités Entomologiques 9: 4–16.

Powell GS, Saxton NA, Pacheco YM, Stanger-Hall KF, Martin GJ, Kusy D, Silveira LFL, Bocak L, Branham MA, Bybee SM. (2022) Beetle bioluminescence outshines extant aerial predators. Proceedings of the Royal Society B: Biological Sciences 289(1979): 20220821. 10.1098/rspb.2022.0821 PubMed DOI PMC

Quintino HYS. (2017) Análise filogenética da subfamília Mastinocerinae LeConte, 1881 (Insecta, Coleoptera, Phengodidae). PhD Thesis, Universidade de São Paulo, São Paulo, Brazil.

Raj JS. (1957) An undescribed luminous beetle larva from south India. Journal of the Bombay Natural History Society 54: 788–789.

Ridley HN. (1934) Further observations made in Singapore, upon Geckos and distasteful moths and upon a luminous Coleopterous, probably Rhagophthalmid, larva. Proceedings of the Royal Entomological Society of London 9: 58–60.

Rosa SP, Costa C, Kramp K, Kundrata R. (2020) Hidden diversity in the Brazilian Atlantic rainforest: The discovery of Jurasaidae, a new beetle family (Coleoptera, Elateroidea) with neotenic females. Scientific Reports 10(1): 1544. 10.1038/s41598-020-58416-6 PubMed DOI PMC

Roza AS. (2020) Notes on the morphology and distribution of the Himalayan genus Pseudothilmanus, including the first record of the genus from Nepal (Coleoptera: Rhagophthalmidae). Transactions of the American Entomological Society 146(2): 421–426. 10.3157/061.146.0208 DOI

Roza AS. (2022) Phylogeny and taxonomy of Phengodidae (Coleoptera: Elateroidea). PhD Thesis, Universidade Federal do Rio de Janeiro, Museu Nacional, Brazil, 1–173.

Roza AS, Mermudes JRM. (2019) New genus and two new species of railroad-worm beetles from Brazil, with a discussion on asymmetry of aedeagus in the family (Coleoptera: Phengodidae). Annales Zoologici 69(4): 805–816. 10.3161/00034541ANZ2019.69.4.012 DOI

Roza AS, Mermudes JRM. (2020) A new genus of railroad-worm beetles from the Atlantic Rainforest from Brazil (Coleoptera: Phengodidae, Mastinocerinae). Papéis Avulsos de Zoologia 60: e202060(s.i.).10. 10.11606/1807-0205/2020.60.special-issue.10 DOI

Roza AS, Quintino HYS, Mermudes JRM, Silveira LFL. (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae). Zootaxa 4306(4): 501–523. 10.11646/zootaxa.4306.4.3 DOI

Roza AS, Mermudes JRM, Silveira LFL. (2018) New species and rediagnosis of Akamboja, and a new record for A.minimum (Coleoptera: Phengodidae, Mastinocerinae). Journal of Natural History 52(45–46): 2935–2947. 10.1080/00222933.2018.1559958 DOI

Rüschkamp F. (1920) Zur biologie der Drilidae und Micromalthidae (Ins. Col.). Biologisches Zentralblatt 40: 376–389.

Sagegami-Oba R, Takahashi N, Oba Y. (2007) The evolutionary process of bioluminescence and aposematism in cantharoid beetles (Coleoptera: Elateroidea) inferred by the analysis of 18S ribosomal DNA. Gene 400(1–2): 104–113. 10.1016/j.gene.2007.06.004 PubMed DOI

Satô F, Kawashima I. (2003) Preliminary report of the genus Stenocladius (Coleoptera, Lampyridae) from Tokashiki-jima Is. and Iheya-jima Is., Okinawa Isls., Southwest Japan. Coleopterists’. News 141: 9–12.

Schaum H. (1850) Bericht über die Leistungen in der Entomologie während des Jahres 1849. Archiv für Naturgeschichte 16: 139–250.

Seri NA, Rahman AA. (2021) Fireflies in South East Asia: Species diversity, distribution, and habitat (2015–2021). Pertanika Journals Tropical Agricultural Science 44: 713–743. 10.47836/pjtas.44.4.02 DOI

Sharp D. (1899) Insects. Part II. Hymenoptera continued (Tubulifera and Aculeata), Coleoptera, Strepsiptera, Lepidoptera, Diptera, Aphaniptera, Thysanoptera, Hemiptera, Anoplura. Macmillan & Co., London, 626 pp. [reprinted 1901 and 1909]

Sheffield NC, Song H, Cameron SL, Whiting MF. (2008) A comparative analysis of mitochondrial genomes in Coleoptera (Arthropoda: Insecta) and genome descriptions of six new beetles. Molecular Biology and Evolution 25(11): 2499–2509. 10.1093/molbev/msn198 PubMed DOI PMC

Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X. (2012) Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research 37: 155–163. 10.1016/j.cretres.2012.03.014 DOI

Sivinski J. (1981) The nature and possible functions of luminescence in Coleoptera larvae. The Coleopterists Bulletin 35: 167–179.

Sivinski JM, Lloyd JE, Beshers SN, Davis LR, Sivinski RG, Sullivan RT, Petersson E. (1998) A natural history of Pleotomodesneedhami Green (Coleoptera: Lampyridae): A firefly symbiont of ants. The Coleopterists Bulletin 52: 23–30.

Stanger-Hall KF, Lloyd JE, Hillis DM. (2007) Phylogeny of North American fireflies (Coleoptera: Lampyridae): Implications for the evolution of light signals. Molecular Phylogenetics and Evolution 45(1): 33–49. 10.1016/j.ympev.2007.05.013 PubMed DOI

Stanger-Hall KF, Sander Lower SE, Lindberg L, Hopkins A, Pallansch J, Hall DW. (2018) The evolution of sexual signal modes and associated sensor morphology in fireflies (Lampyridae, Coleoptera). Proceedings. Biological Sciences 285(1871): 20172384. 10.1098/rspb.2017.2384 PubMed DOI PMC

Suzuki H. (1997) Molecular phylogenetic studies of Japanese fireflies and their mating systems (Coleoptera: Cantharoidea). Tokyo Metropolitan University Bulletin of Natural History 3: 1–53.

Suzuki H, Kobayashi Y. (2009) Embryogenesis of firefly: Phylogenetic implication of the family Rhagophthalmidae. In: Napompeth B. (Ed.) International Symposium on Diversity and Conservation of Fireflies.Queen Sirikit Botanic Garden, Chiang Mai, Thailand, 30–35.

Tan JCH. (2018) Fireflies of Pulau Ubin, Singapore. Nature in Singapore 11: 127–138.

Timmermans MJTN, Vogler AP. (2012) Phylogenetically informative rearrangements in mitochondrial genomes of Coleoptera, and monophyly of aquatic elateriform beetles (Dryopoidea). Molecular Phylogenetics and Evolution 63(2): 299–304. 10.1016/j.ympev.2011.12.021 PubMed DOI

Ugarova NN, Brovko LY. (2002) Protein structure and bioluminescent spectra for firefly bioluminescence. Luminescence 17(5): 321–330. 10.1002/bio.688 PubMed DOI

Vega-Badillo V, Zaragoza-Caballero S. (2019) New species of the genus Phengodes (Phengodella) (Coleoptera: Phengodidae) and a key to phengodids from Belize. Revista Mexicana de Biodiversidad 90: e902863. 10.22201/ib.20078706e.2019.90.2863 DOI

Vega-Badillo V, Zaragoza-Caballero S, Ivie MA. (2020) A new genus of Phengodidae (Coleoptera) from the Neotropical Region. Papéis Avulsos de Zoologia 60: e202060(s.i.).06. 10.11606/1807-0205/2020.60.special-issue.06 DOI

Vega-Badillo V, Morrone JJ, Zaragoza-Caballero S. (2021a) Revision of the genus Cenophengus LeConte, 1881 (Coleoptera, Phengodidae), with the description of four new species, new geographic records and a new synonymy. ZooKeys 1068: 73–148. 10.3897/zookeys.1068.70295 PubMed DOI PMC

Vega-Badillo V, Zaragoza-Caballero S, Rios-Ibarra JJ. (2021b) New species of the genus Cenophengus LeConte 1881 (Coleoptera; Phengodidae) from Mexico and Guatemala. Zootaxa 5023(2): 223–238. 10.11646/zootaxa.5023.2.3 PubMed DOI

Villiers A. (1958) Maurice Pic †. Entomologische Blätter 54: 3–4.

Viviani VR. (2002) The origin, diversity, and structure-function relationships of insect luciferases. Cellular and Molecular Life Sciences 59(11): 1833–1850. 10.1007/PL00012509 PubMed DOI PMC

Viviani VR, Bechara EJH. (1993) Biophysical and biochemical aspects of phengodid (railroad-worm) bioluminescence. Photochemistry and Photobiology 58(4): 615–622. 10.1111/j.1751-1097.1993.tb04941.x DOI

Viviani VR, Bechara EJH. (1997) Bioluminescence and biological aspects of Brazilian railroad-worms (Coleoptera: Phengodidae). Annals of the Entomological Society of America 90(3): 389–398. 10.1093/aesa/90.3.389 DOI

Viviani VR, Ohmiya Y. (2000) Bioluminescence color determinants of Phrixothrix railroad-worm luciferases: Chimeric luciferases, site-directed mutagenesis of Arg 215 and guanidine effect. Photochemistry and Photobiology 72(2): 267–271. 10.1562/0031-8655(2000)072<0267:BCDOPR>2.0.CO;2 PubMed DOI

Viviani VR, Bechara EJH, Ohmiya Y. (1999) Cloning, sequence analysis, and expression of active Phrixothrix railroad-worms luciferases: Relationship between bioluminescence spectra and primary structures. Biochemistry 38(26): 8271–8279. 10.1021/bi9900830 PubMed DOI

Viviani VR, Uchida A, Suenaga N, Ryufuku M, Ohmiya Y. (2001) Thr226 is a key residue for bioluminescence spectra determination in beetle luciferases. Biochemical and Biophysical Research Communications 280(5): 1286–1291. 10.1006/bbrc.2001.4254 PubMed DOI

Viviani VR, Uchida A, Viviani W, Ohmiya Y. (2002) The influence of Ala243 (Gly247), Arg215 and Thr226 (Asn230) on the bioluminescence spectra and pH-sensitivity of railroad worm, click beetle and firefly luciferases. Photochemistry and Photobiology 76(5): 538–544. 10.1562/0031-8655(2002)076<0538:TIOAGA>2.0.CO;2 PubMed DOI

Voris HK. (2000) Maps of Pleistocene sea levels in Southeast Asia: Shorelines, river systems and time durations. Journal of Biogeography 27(5): 1153–1167. 10.1046/j.1365-2699.2000.00489.x DOI

Wallace AR. (1863) On the Physical Geography of the Malay Archipelago. Royal Geographical Society 7: 205–212. 10.2307/1798448 DOI

Wang K, Hong W, Jiao H, Zhao H. (2017) Transcriptome sequencing and phylogenetic analysis of four species of luminescent beetles. Scientific Reports 7(1): 1814. 10.1038/s41598-017-01835-9 PubMed DOI PMC

Waterhouse CO. (1890) Aid to the identification of insects. Vol. 2. EW Janson, London, 1–29.

Westwood JO. (1849) Dodecatoma. In: Guérin-Méneville FE (Ed.) Species et iconographie générique des animaux articulés ou représentation des genres, avec leur description et celle de toutes les espèces de cette grande division du règne animal. Ouvrage formant une série de Monographies complètes. Insectes coléoptères. Livraison 4. Bureau de la Revue Zoologique et Magasin de Zoologie, Paris, France, No. 16, 1 unnumbered page [1843].

Wijekoon WMCD, Wegiriya HCE, Bogahawatta CNL. (2016) Systematic revision of the repository collection of Canthoroidea in the Department of National Museums, Colombo, Sri Lanka (Coleoptera: Antharidae, Lampyridae, Lycidae, Rhagophthalmidae). Ceylon Journal of Science 45(1): 67–74. 10.4038/cjs.v45i1.7365 DOI

Winkler A. (1925) Catalogus Coleopterorum regionis palaearcticae. I. A. Caraboidea, B. Palpicornia, Staphylinoidea, C. Diversicornia. Pars 5 [pp. 497–624]. Albert Winkler, Wien, 1698 pp.

Wittmer W. (1938) 3. Beitrag zur Kenntnis der Indo-Malayischen Malacodermata (Col.). Treubia 16: 301–306.

Wittmer W. (1939) 5. Beitrag zur Kenntnis der Indo-Malayischen Malacodermata (Col.). Treubia 17: 21–32.

Wittmer W. (1941) Malacodermata von den Philippinen aus der Sammlung des zoologischen Museums in Amsterdam. Philippine Journal of Science 74: 195–228.

Wittmer W. (1944) Catalogue des Drilidae E. Oliv. (Coleoptera-Malacodermata). Revista de la Sociedad Entomológica Argentina XII: 203–221.

Wittmer W. (1948) Supplement au catalogue des Drilidae E. Oliv. (Col. Malacodermata). Revista de la Sociedad Entomológica Argentina XIV: 115–116.

Wittmer W. (1979) Zur Kenntnis der Lampyridae der orientalischen Region (Coleoptera). Entomologische Arbeiten aus dem Museum G. Frey 28: 83–92.

Wittmer W. (1995) Lampyridae und Omethidae aus dem indo-malaiischen Faunengebiete (Coleoptera). Mitteilungen der Entomologischen Gesellschaft Basel 45: 106–117.

Wittmer W. (1997) Neue Rhagophthalmidae und Phengodidae (Coleoptera). Mitteilungen der Schweizerische Entomologische Gesellschaft 70: 257–263.

Wittmer W, Ohba N. (1994) Neue Rhagophthalmidae (Coleoptera) aus China und benachbarten Ländern. Japanese Journal of Entomology 62: 341–355.

Wu CF. (1937) Catalogus Insectorum Sinensium. Vol. III. Fan Memorial Institute of Biology, Beijing, China, 1312 pp.

Yiu V. (2011a) Principles of Chinese nomenclature of fireflies and Chinese names of Hong Kong fireflies. Insect News 2: 13–16. [Hong Kong Entomological Society Newsletter]

Yiu V. (2011b) Observations on the luminescence configurations of eight firefly genera and their immature stages. Hong Kong Entomological Bulletin 3: 20–30.

Yiu V. (2012) Fireflies of Hong Kong. Popular Entomology Book Series, Volume 7. Hong Kong Entomological Society, 117 pp.

Yiu V. (2013) A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 1. Lampyrid 3: 100–126.

Yiu V. (2017) A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2. Lampyrid 4: 59–111. [67–119]

Yiu V, Jeng ML. (2018) Oculogryphuschenghoiyanae sp. n. (Coleoptera, Lampyridae): A new ototretine firefly from Hong Kong with descriptions of its bioluminescent behavior and ultraviolet-induced fluorescence in females. ZooKeys 739: 65–78. 10.3897/zookeys.739.21502 PubMed DOI PMC

Zaragoza-Caballero S, Hernández CXP. (2014) Sinopsis de la familia Phengodidae (Coleoptera): trenecitos, bigotudos, glow-worms, railroad-worms o besouros trem de ferro. Instituto de Biología, Universidad Nacional Autónoma de México, México, D.F., 1–130. 10.22201/IB.9786070251832E.2014 DOI

Zaragoza-Caballero S, Zurita-García ML. (2015) A preliminary study on the phylogeny of the family Phengodidae (Insecta: Coleoptera). Zootaxa 3947(4): 527–542. 10.11646/zootaxa.3947.4.4 PubMed DOI

Zhang S-Q, Che L-H, Li Y, Liang D, Pang H, Ślipiński A, Zhang P. (2018) Evolutionary history of Coleoptera revealed by extensive sampling of genes and species. Nature Communications 9(1): 205. 10.1038/s41467-017-02644-4 PubMed DOI PMC

Zhang R, He J, Dong Z, Liu G, Yin Y, Zhang X, Li Q, Ren Y, Yang Y, Liu W, Chen X, Xia W, Duan K, Hao F, Lin Z, Yang J, Chang Z, Zhao R, Wan W, Lu S, Peng Y, Ge S, Wang W, Li X. (2020) Genomic and experimental data provide new insights into luciferin biosynthesis and bioluminescence evolution in fireflies. Scientific Reports 10(1): 15882. 10.1038/s41598-020-72900-z PubMed DOI PMC

Discovery of a new soft-bodied click-beetle genus from Namibia with a unique morphology leads to a modified diagnosis of Drilini (Coleoptera, Elateridae)

A light in the dark: a mid-Cretaceous bioluminescent firefly with specialized antennal sensory organs

A new species of Rhagophthalmus Motschulsky, 1854 (Coleoptera, Rhagophthalmidae) from Laos represents the smallest known member of the genus