Based on recent achievements in phylogenetic studies of the Brassicaceae, a novel infrafamilial classification is proposed that includes major improvements at the subfamilial and supertribal levels. Herein, the family is subdivided into two subfamilies, Aethionemoideae (subfam. nov.) and Brassicoideae. The Brassicoideae, with 57 of the 58 tribes of Brassicaceae, are further partitioned into five supertribes, including the previously recognized Brassicodae and the newly established Arabodae, Camelinodae, Heliophilodae, and Hesperodae. Additional tribus-level contributions include descriptions of the newly recognized Arabidopsideae, Asperuginoideae, Hemilophieae, Schrenkielleae, and resurrection of the Chamireae and Subularieae. Further detailed comments on 17 tribes in need of clarifications are provided.

Central European Institute of Technology and Faculty of Science Masaryk University Kamenice 5 62500 Brno Czech Republic Masaryk University Brno Czech Republic

Department of Biodiversity and Plant Systematics Centre for Organismal Studies Heidelberg University Im Neuenheimer Feld 345 D 69120 Heidelberg Germany Heidelberg University Heidelberg Germany

Department of Biology Botany University of Osnabrück Barbarastraße 11 49076 Osnabrück Germany University of Osnabrück Osnabrück Germany

Department of Biology New Mexico State University P O Box 30001 MSC 3AF Las Cruces NM 88003 USA New Mexico State University Las Cruces United States of America

Institute of Biology Leiden Leiden University Sylviusweg 72 2333 BE Leiden Netherlands Leiden University Leiden Netherlands

Missouri Botanical Garden 4344 Shaw Boulevard St Louis Missouri 63110 USA Missouri Botanical Garden St Louis United States of America

Naturalis Biodiversity Center Darwinweg 2 2333 CR Leiden Netherlands Naturalis Biodiversity Center Leiden Netherlands

South Siberian Botanical Garden Altai State University Lenin Ave 61 656049 Barnaul Russia Altai State University Barnaul Russia

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