Theories explain the presence of fears and specific phobias elicited by animals in contemporary WEIRD (Western, educated, industrialized, rich, and democratic) populations by their evolutionary past in Africa. Nevertheless, empirical data about fears of animals in the Cradle of Humankind are still fragmentary. To fill this gap, we examined which local animals are perceived as the most frightening by Somali people, who inhabit a markedly similar environment and the region where humans have evolved. We asked 236 raters to rank 42 stimuli according to their elicited fear. The stimuli were standardized pictures of species representing the local fauna. The results showed that the most frightening animals were snakes, scorpions, the centipede, and large carnivores (cheetahs and hyenas). These were followed up by lizards and spiders. Unlike in Europe, spiders represent less salient stimuli than scorpions for Somali respondents in this study. This conforms to the hypothesis suggesting that fear of spiders was extended or redirected from other chelicerates.

Amoud University Borama Somaliland

Department of Zoology Faculty of Science Charles University Viničná 7 128 43 Prague 2 Czech Republic

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Abay GY, Bauer H, Gebrihiwot K, Deckers J. Peri-urban spotted hyena (Crocuta crocuta) in Northern Ethiopia: diet, economic impact, and abundance. Eur J Wildlife Res. 2011;57(4):759–765. doi: 10.1007/s10344-010-0484-8. DOI

Baynes-Rock M. Human perceptual and phobic biases for snakes: a review of the experimental evidence. Anthrozoös. 2017;30(1):5–18. doi: 10.1080/08927936.2017.1270584. DOI

Bertels J, De Heering A, Bourguignon M, Cleeremans A, Destrebecqz A. What determines the neural response to snakes in the infant brain? A systematic comparison of color and grayscale stimuli. Front Psychol. 2023;14:1027872. doi: 10.3389/fpsyg.2023.1027872. PubMed DOI PMC

Casewell NR, Jackson TN, Laustsen AH, Sunagar K. Causes and consequences of snake venom variation. Trends Pharmacol Sci. 2020;41(8):570–581. doi: 10.1016/j.tips.2020.05.006. PubMed DOI PMC

Castelló JR. Bovids of the world: antelopes, gazelles, cattle, goats, sheep, and relatives. Princeton University Press, Princeton & Oxford; 2016.

Ceríaco LM. Human attitudes towards herpetofauna: the influence of folklore and negative values on the conservation of amphibians and reptiles in Portugal. J Ethnobiol Ethnomed. 2012;8(1):1–13. doi: 10.1186/1746-4269-8-8. PubMed DOI PMC

Ceríaco LM, Marques MP, Madeira NC, Vila-Viçosa CM, Mendes P. Folklore and traditional ecological knowledge of geckos in Southern Portugal: implications for conservation and science. J Ethnobiol Ethnomed. 2011;7(1):1–10. doi: 10.1186/1746-4269-7-26. PubMed DOI PMC

Chippaux JP. Snake venoms and envenomations. Malabar: Krieger Publishers; 2006.

Chippaux JP, Goyfon M. Epidemiology of scorpionism: a global appraisal. Acta Trop. 2008;107:71–79. doi: 10.1016/j.actatropica.2008.05.021. PubMed DOI

Chomba C, Senzota R, Chabwela H, Mwitwa J, Nyirenda V. Patterns of human-wildlife conflicts in Zambia, causes, consequences and management responses. J Ecol Nat Environ. 2012;4(12):303–313. doi: 10.5897/JENE12.029. DOI

Coelho CM, Purkis H. The origins of specific phobias: influential theories and current perspectives. Rev Gen Psychol. 2009;13(4):335–348. doi: 10.1037/a0017759. DOI

Coelho CM, Gonçalves-Bradley D, Zsido AN. Who worries about specific phobias?–A population-based study of risk factors. J Psych Res. 2020;126:67–72. doi: 10.1016/j.jpsychires.2020.05.001. PubMed DOI

Dawkins R, Wong Y. The ancestor’s tale: a pilgrimage to the dawn of life. UK: Hachette; 2016.

de Pinho JR, Grilo C, Boone RB, Galvin KA, Snodgrass JG. Influence of aesthetic appreciation of wildlife species on attitudes towards their conservation in Kenyan agropastoralist communities. PloS one. 2014;9(2):e88842. doi: 10.1371/journal.pone.0088842. PubMed DOI PMC

del Hoyo J, Elliott A, Christie D. Handbook of the birds of the world. Barcelona: Lynx editions; 1996.

Dressel S, Sandström C, Ericsson G. A meta-analysis of studies on attitudes toward bears and wolves across Europe 1976-2012. Conserv Biol. 2015;29(2):565–574. doi: 10.1111/cobi.12420. PubMed DOI

Dunham KM, Ghiurghi A, Cumbi R, Urbano F. Human–wildlife conflict in Mozambique: a national perspective, with emphasis on wildlife attacks on humans. Oryx. 2010;44(2):185–193. doi: 10.1017/S003060530999086X. DOI

Elliot DG. List of mammals from Somaliland obtained by the Museum’s East African expedition. Fieldiana Zool. 1897;1(6):1–54.

Evangelista PH, Mohamed AM, Hussein IA, Saied AH, Mohammed AH, Young NE. Integrating indigenous local knowledge and species distribution modelling to detect wildlife in Somaliland. Ecosphere. 2018;9(3):e02134. doi: 10.1002/ecs2.2134. DOI

Evans KD, Swedell L, Chowdhury S. Respect for males amid suppression of females: selective use of aggression and fitness correlates in the male-dominated society of hamadryas baboons. J Hum Evol. 2022;165:103151. doi: 10.1016/j.jhevol.2022.103151. PubMed DOI

Fabio RA, Caprì T. Automatic and controlled attentional capture by threatening stimuli. Heliyon. 2019;5(5):e01752. doi: 10.1016/j.heliyon.2019.e01752. PubMed DOI PMC

Forde A, Jacobsen A, Dugon MM, Healy K. Scorpion species with smaller body sizes and narrower chelae have the highest venom potency. Toxins. 2022;14(3):219. doi: 10.3390/toxins14030219. PubMed DOI PMC

Fredrikson M, Annas P, Fischer H, Wik G. Gender and age differences in the prevalence of specific fears and phobias. Behav Res Ther. 1996;34(1):33–39. doi: 10.1016/0005-7967(95)00048-3. PubMed DOI

Frynta D, Lišková S, Bültmann S, Burda H. Being attractive brings advantages: the case of parrot species in captivity. PloS one. 2010;5(9):e12568. doi: 10.1371/journal.pone.0012568. PubMed DOI PMC

Frynta D, Marešová J, Řeháková-Petrů M, Šklíba J, Šumbera R, Krása A. Cross-cultural agreement in perception of animal beauty: boid snakes viewed by people from five continents. Hum Ecol. 2011;39(6):829–834. doi: 10.1007/s10745-011-9447-2. DOI

Frynta D, Šimková O, Lišková S, Landová E. Mammalian collection on Noah’s ark: the effects of beauty, brain and body size. PloS one. 2013;8(5):e63110. doi: 10.1371/journal.pone.0063110. PubMed DOI PMC

Frynta D, Janovcová M, Štolhoferová I, Peléšková Š, Vobrubová B, Frýdlová P, Skalíková H, Šípek P, Landová E. Emotions triggered by live arthropods shed light on spider phobia. Sci Rep. 2021;11(1):1–10. doi: 10.1038/s41598-021-01325-z. PubMed DOI PMC

Gamer M, Lemon J, Singh P (2022) Package ‘irr’. https://cran.r-project.org/web/packages/irr/irr.pdf. Accessed 14 Nov 2022

Gebresenbet F, Bauer H, Vadjunec JM, Papeş M. Beyond the numbers: human attitudes and conflict with lions (Panthera leo) in and around Gambella National Park. Ethiopia. PloS one. 2018;13(9):e0204320. doi: 10.1371/journal.pone.0204320. PubMed DOI PMC

Groucutt HS, Petraglia MD, Bailey G, Scerri EM, Parton A, Clark-Balzan L, Jennings RP, Lewis L, Blinkhorn J, Drake NA, Breeze PS. Rethinking the dispersal of Homo sapiens out of Africa. Evol Anthropol. 2015;24(4):149–164. doi: 10.1002/evan.21455. PubMed DOI PMC

Harris RJ, Nekaris KAI, Fry BG. Monkeying around with venom: an increased resistance to α-neurotoxins supports an evolutionary arms race between Afro-Asian primates and sympatric cobras. BMC Biol. 2021;19:1–13. doi: 10.1186/s12915-021-01195-x. PubMed DOI PMC

Hauke TJ, Herzig V. Dangerous arachnids—fake news or reality? Toxicon. 2017;138:173–183. doi: 10.1016/j.toxicon.2017.08.024. PubMed DOI

Hayward MW, Henschel P, O'Brien J, Hofmeyr M, Balme G, Kerley GI. Prey preferences of the leopard (Panthera pardus) J Zool. 2006;270(2):298–313. doi: 10.1111/j.1469-7998.2006.00139.x. DOI

Hill CM. Crop-raiding by wild vertebrates: the farmer’s perspective in an agricultural community in western Uganda. Int J Pest Manage. 1997;43(1):77–84. doi: 10.1080/096708797229022. DOI

Isbell LA. Snakes as agents of evolutionary change in primate brains. J Hum Evol. 2006;51(1):1–35. doi: 10.1016/j.jhevol.2005.12.012. PubMed DOI

Isbell LA. The fruit, the tree, and the serpent: why we see so well. Harvard University Press; 2009.

Isbell LA, Etting SF. Scales drive detection, attention, and memory of snakes in wild vervet monkeys (Chlorocebus pygerythrus) Primates. 2017;58(1):121–129. doi: 10.1007/s10329-016-0562-y. PubMed DOI

Isbister GK, Bawaskar HS. Scorpion envenomation. New Engl J Med. 2014;371(5):457–463. doi: 10.1056/NEJMra1401108. PubMed DOI

Janovcová M, Rádlová S, Polák J, Sedláčková K, Peléšková Š, Žampachová B, Frynta D, Landová E. Human attitude toward reptiles: a relationship between fear, disgust, and aesthetic preferences. Animals. 2019;9(5):238. doi: 10.3390/ani9050238. PubMed DOI PMC

Johansson M, Ferreira IA, Støen OG, Frank J, Flykt A. Targeting human fear of large carnivores—many ideas but few known effects. Biol Conserv. 2016;201:261–269. doi: 10.1016/j.biocon.2016.07.010. DOI

Kaltenborn BRP, Bjerke T, Nyahongo J. Living with problem animals—self-reported fear of potentially dangerous species in the Serengeti Region, Tanzania. Hum Dimens Wildl. 2006;11(6):397–409. doi: 10.1080/10871200600984323. DOI

Kanga EM, Ogutu JO, Piepho HP, Olff H. Human–hippo conflicts in Kenya during 1997–2008: vulnerability of a megaherbivore to anthropogenic land use changes. J Land Use Sci. 2012;7(4):395–406. doi: 10.1080/1747423X.2011.590235. DOI

Kansky R, Kidd M, Knight AT. Meta-analysis of attitudes toward damage-causing mammalian wildlife. Conserv Biol. 2014;28(4):924–938. doi: 10.1111/cobi.12275. PubMed DOI PMC

Kisker J, Lange L, Flinkenflügel K, Kaup M, Labersweiler N, Tetenborg F, Ott P, Gundler C, Gruber T, Osinsky R, Schöne B. Authentic fear responses in virtual reality: a mobile EEG study on affective, behavioral and electrophysiological correlates of fear. Front Virtual Real. 2021;2:716318. doi: 10.3389/frvir.2021.716318. DOI

Koch MD, O’Neill HK, Sawchuk CN, Connolly K. Domain-specific and generalized disgust sensitivity in blood-injection-injury phobia:: the application of behavioral approach/avoidance tasks. J Anxiety Disord. 2002;16(5):511–527. doi: 10.1016/S0887-6185(02)00170-6. PubMed DOI

Kovařík F, Lowe G, Elmi HSA, Šťáhlavský F. Scorpions of the horn of Africa (Arachnida: Scorpiones). Part XXI. Parabuthus (Buthidae)(Part II), with description of five new species from Somaliland and Ethiopia. Euscorpius. 2019;2019(290):1–63. doi: 10.18590/euscorpius.2019.vol2019.iss290.1. DOI

Landová E, Marešová J, Šimková O, Cikánová V, Frynta D. Human responses to live snakes and their photographs: evaluation of beauty and fear of the king snakes. J Environ Psychol. 2012;32(1):69–77. doi: 10.1016/j.jenvp.2011.10.005. DOI

Landová E, Bakhshaliyeva N, Janovcová M, Peléšková Š, Suleymanova M, Polák J, Guliev A, Frynta D. Association between fear and beauty evaluation of snakes: cross-cultural findings. Front Psychol. 2018;9:333. doi: 10.3389/fpsyg.2018.00333. PubMed DOI PMC

Landová E, Poláková P, Rádlová S, Janovcová M, Bobek M, Frynta D. Beauty ranking of mammalian species kept in the Prague Zoo: does beauty of animals increase the respondents’ willingness to protect them? Sci Nat. 2018;105(11):1–14. doi: 10.1007/s00114-018-1596-3. PubMed DOI

Landová E, Janovcová M, Štolhoferová I, Rádlová S, Frýdlová P, Sedláčková K, Frynta D. Specificity of spiders among fear-and disgust-eliciting arthropods: spiders are special, but phobics not so much. PloS one. 2021;16(9):e0257726. doi: 10.1371/journal.pone.0257726. PubMed DOI PMC

Landová E, Peléšková Š, Sedláčková K, Janovcová M, Polák J, Rádlová S, Vobrubová B, Frynta D. Venomous snakes elicit stronger fear than nonvenomous ones: psychophysiological response to snake images. Plos one. 2020;15(8):e0236999. doi: 10.1371/journal.pone.0236999. PubMed DOI PMC

Landová E, Rádlová S, Pidnebesna A, Tomeček D, Janovcová M, Peléšková Š, Sedláčková K, Štolhoferová I, Polák J, Hlinka J, Frynta D. Towards a reliable detection of arachnophobia: subjective, behavioral, and neurophysiological measures of fear response. Front. Psych. 2023;14:1025. doi: 10.3389/fpsyt.2023.1196785. PubMed DOI PMC

Lišková S, Frynta D. What determines bird beauty in human eyes? Anthrozoös. 2013;26(1):27–41. doi: 10.2752/175303713X13534238631399. DOI

Lišková S, Landová E, Frynta D. Human preferences for colorful birds: vivid colors or pattern? Evol Psychol. 2015;13(2):147470491501300203. doi: 10.1177/147470491501300203. PubMed DOI

Marešová J, Krása A, Frynta D. We all appreciate the same animals: cross-cultural comparison of human aesthetic preferences for snake species in Papua New Guinea and Europe. Ethology. 2009;115(4):297–300. doi: 10.1111/j.1439-0310.2009.01620.x. DOI

Marešová J, Landová E, Frynta D. What makes some species of milk snakes more attractive to humans than others? Theory Biosci. 2009;128(4):227–235. doi: 10.1007/s12064-009-0075-y. PubMed DOI

Matema S, Andersson JA. Why are lions killing us? Human–wildlife conflict and social discontent in Mbire District, northern Zimbabwe. J Mod Afr Stud. 2015;53(1):93–120. doi: 10.1017/S0022278X14000664. DOI

Mavridis IN, Meliou M, Pyrgelis ES. Clinical consequences of centipede bite: is it neurotoxic? World J Neurol. 2016;6(2):23–29. doi: 10.5316/wjn.v6.i2.23. DOI

McDougall I, Brown FH, Fleagle JG. Stratigraphic placement and age of modern humans from Kibish. Ethiopia. Nature. 2005;433(7027):733–736. doi: 10.1038/nature03258. PubMed DOI

Mills MGL, Mills M. Kalahari cheetahs: adaptations to an arid region. Oxford: Oxford University Press; 2017.

Muris P, du Plessis M, Loxton H. Origins of common fears in South African children. J Anxiety Disord. 2008;22(8):1510–1515. doi: 10.1016/j.janxdis.2008.03.004. PubMed DOI

Öhman A, Flykt A, Esteves F. Emotion drives attention: detecting the snake in the grass. J Exp Psychol. 2001;130(3):466. doi: 10.1037/0096-3445.130.3.466. PubMed DOI

Öhman A, Mineka S. Fears, phobias, and preparedness: toward an evolved module of fear and fear learning. Psychol Rev. 2001;108(3):483. doi: 10.1037//0033-295X.108.3.483. PubMed DOI

Öhman A, Soares SC, Juth P, Lindström B, Esteves F. Evolutionary derived modulations of attention to two common fear stimuli: serpents and hostile humans. J Cogn Psychol. 2012;24(1):17–32. doi: 10.1080/20445911.2011.629603. DOI

Ombati R, Luo L, Yang S, Lai R. Centipede envenomation: clinical importance and the underlying molecular mechanisms. Toxicon. 2018;154:60–68. doi: 10.1016/j.toxicon.2018.09.008. PubMed DOI

Onyishi IE, Nwonyi SK, Pazda A, Prokop P. Attitudes and behaviour toward snakes on the part of Igbo people in southeastern Nigeria. Sci Total Environ. 2021;763:143045. doi: 10.1016/j.scitotenv.2020.143045. PubMed DOI

Packer C, Ikanda D, Kissui B, Kushnir H. Lion attacks on humans in Tanzania. Nature. 2005;436:927–928. doi: 10.1038/436927a. PubMed DOI

Packer C, Swanson A, Ikanda D, Kushnir H. Fear of darkness, the full moon and the nocturnal ecology of African lions. PloS one. 2011;6(7):e22285. doi: 10.1371/journal.pone.0022285. PubMed DOI PMC

Pohlert T (2014) The Pairwise Multiple Comparison of Mean Ranks Package (PMCMR). R Package. https://CRAN.R-project.org/package=PMCMR, (accessed on 24 Sept 2020).

Polák J, Sedláčková K, Nácar D, Landová E, Frynta D. Fear the serpent: a psychometric study of snake phobia. Psych Res. 2016;242:163–168. doi: 10.1016/j.psychres.2016.05.024. PubMed DOI

Polák J, Rádlová S, Janovcová M, Flegr J, Landová E, Frynta D. Scary and nasty beasts: self-reported fear and disgust of common phobic animals. Br J Psychol. 2020;111(2):297–321. doi: 10.1111/bjop.12409. PubMed DOI

Polák J, Sedláčková K, Janovcová M, Peléšková Š, Flegr J, Vobrubová B, Frynta D, Landová E. Measuring fear evoked by the scariest animal: Czech versions of the Spider Questionnaire and Spider Phobia Beliefs Questionnaire. BMC Psychiatry. 2022;22(1):1–13. doi: 10.1186/s12888-021-03672-7. PubMed DOI PMC

Profico A, Di Vincenzo F, Gagliardi L, Piperno M, Manzi G. Filling the gap. Human cranial remains from Gombore II (Melka Kunture, Ethiopia; ca. 850 ka) and the origin of Homo heidelbergensis. J Anthropol Sci. 2016;94:1–24. doi: 10.4436/jass.94019. PubMed DOI

Prokop P, Tolarovičová A, Camerik AM, Peterková V. High school students’ attitudes towards spiders: a cross-cultural comparison. Int J Sci Educ. 2010;32(12):1665–1688. doi: 10.1080/09500690903253908. DOI

Prokop P, Fančovičová J, Kučerová A. Aposematic colouration does not explain fear of snakes in humans. J Ethol. 2018;36:35–41. doi: 10.1007/s10164-017-0533-9. DOI

Rádlová S, Janovcová M, Sedláčková K, Polák J, Nácar D, Peléšková Š, Frynta D, Landová E. Snakes represent emotionally salient stimuli that may evoke both fear and disgust. Front Psychol. 2019;10:1085. doi: 10.3389/fpsyg.2019.01085. PubMed DOI PMC

Rádlová S, Polák J, Janovcová M, Sedláčková K, Peléšková Š, Landová E, Frynta D. Emotional reaction to fear-and disgust-evoking snakes: sensitivity and propensity in snake-fearful respondents. Front Psychol. 2020;11:31. doi: 10.3389/fpsyg.2020.00031. PubMed DOI PMC

Reider LB, Mahaffey EM, Barylski B, LoBue V. “It bites!”: The transmission of negative information about snakes and spiders through a naturalistic picture book interaction. Dev Psychol. 2022;58(11):2140–2157. doi: 10.1037/dev0001429. PubMed DOI

R Development Core Team . R: a language and environment for statistical computing. Vienna: R Foundation for Statistical Computing; 2010.

Rito T, Vieira D, Silva M, Conde-Sousa E, Pereira L, Mellars P, Richards MB, Soares P. A dispersal of Homo sapiens from southern to eastern Africa immediately preceded the out-of-Africa migration. Sci Rep. 2019;9(1):1–10. doi: 10.1038/s41598-019-41176-3. PubMed DOI PMC

Rudolfová V, Štolhoferová I, Elmi HSA, Rádlová S, Rexová K, Berti DA, Král D, Sommer D, Landová E, Frýdlová P, Frynta D. Do spiders ride on the fear of scorpions? A cross-cultural eye tracking study. Animals. 2022;12(24):3466. doi: 10.3390/ani12243466. PubMed DOI PMC

Sanitbanez-Lopez CE, Aharon S, Ballesteroset JA et al (2020) Phylogenomics of scorpions reveal a co-diversification of scorpion mammalian predators and mammal-specifc sodium channel toxins. bioRxiv. 10.1101/2020.11.06.372045

Schaffrath S, Prendini L, Predel R. Intraspecific venom variation in southern African scorpion species of the genera Parabuthus, Uroplectes and Opistophthalmus (Scorpiones: Buthidae, Scorpionidae) Toxicon. 2018;144:83–90. doi: 10.1016/j.toxicon.2018.02.004. PubMed DOI

Seligman ME. Phobias and preparedness. Behav Ther. 1971;2(3):307–320. doi: 10.1016/S0005-7894(71)80064-3. DOI

Soga M, Gaston KJ. Extinction of experience: the loss of human-nature interactions. Front Ecol Environ. 2016;14(2):94–101. doi: 10.1002/fee.1225. DOI

Staňková H, Janovcová M, Peléšková Š, Sedláčková K, Landová E, Frynta D. The ultimate list of the most frightening and disgusting animals: negative emotions elicited by animals in Central European respondents. Animals. 2021;11(3):747. doi: 10.3390/ani11030747. PubMed DOI PMC

StatSoft Inc (2010) Statistica (data analysis software system), version 9.1. Available online at http://www.statsoft.com. Accessed 6 Apr 2010

Stringer C. The origin and evolution of Homo sapiens. Philos T Roy Soc B. 2016;371(1698):20150237. doi: 10.1098/rstb.2015.0237. PubMed DOI PMC

Treves A, Naughton-Treves L. Risk and opportunity for humans coexisting with large carnivores. J Hum Evol. 1999;36(3):275–282. doi: 10.1006/jhev.1998.0268. PubMed DOI

Undheim EA, Jenner RA. Phylogenetic analyses suggest centipede venom arsenals were repeatedly stocked by horizontal gene transfer. Nat Commun. 2021;12(1):1–14. doi: 10.1038/s41467-021-21093-8. PubMed DOI PMC

Valenta J. Venomous snakes – envenoming, therapy. New York: Nova Science Publishers; 2010.

Van Le Q, Isbell LA, Matsumoto J, Nguyen M, Hori E, Maior RS, Tomazc C, Trana AH, Onoa T, Nishijo H. Pulvinar neurons reveal neurobiological evidence of past selection for rapid detection of snakes. P Natl Acad Sci USA. 2013;110(47):19000–19005. doi: 10.1073/pnas.1312648110. PubMed DOI PMC

Van Strien JW, Isbell LA. Snake scales, partial exposure, and the Snake Detection Theory: a human event-related potentials study. Sci Rep. 2017;7(1):1–9. doi: 10.1038/srep46331. PubMed DOI PMC

Vetter RS, Draney ML, Brown CA, Trumble JT, Gouge DH, Hinkle NC, Pace-Schott EF. Spider fear versus scorpion fear in undergraduate students at five American Universities. Am Entomol. 2018;64(2):79–82. doi: 10.1093/ae/tmy030. DOI

Ward MJ, Ellsworth SA, Nystrom GS. A global accounting of medically signifcant scorpions: Epidemiology, major toxins, and comparative resources in harmless counterparts. Toxicon. 2018;151:137–155. doi: 10.1016/j.toxicon.2018.07.007. PubMed DOI

Weiss L, Brandl P, Frynta D. Fear reactions to snakes in naïve mouse lemurs and pig-tailed macaques. Primates. 2015;56(3):279–284. doi: 10.1007/s10329-015-0473-3. PubMed DOI

White TD, Asfaw B, DeGusta D, Gilbert H, Richards GD, Suwa G, Howell FC. Pleistocene homo sapiens from middle awash, ethiopia. Nature. 2003;423(6941):742–747. doi: 10.1038/nature01669. PubMed DOI

Wilson DE, Mittermeier RA. Handbook of the mammals of the world. Barcelona: Lynx Edicions; 2009.

Yirga G, Leirs H, De Iongh HH, Asmelash T, Gebrehiwot K, Deckers J, Bauer H. Spotted hyena (Crocuta crocuta) concentrate around urban waste dumps across Tigray, northern Ethiopia. Wildl Res. 2015;42(7):563–569. doi: 10.1071/WR14228. DOI

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